Conchology, from

Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...

κόγχος (''kónkhos''), meaning " cockle", and

-logy ''-logy'' is a suffix in the English language, used with words originally adapted from Ancient Greek ending in ('). The earliest English examples were anglicizations of the French '' -logie'', which was in turn inherited from the Latin '' -l ...

from λόγος (''lógos''), meaning "study", is the study of

mollusc shell The mollusc (or mollusk) shell is typically a calcareous exoskeleton which encloses, supports and protects the soft parts of an animal in the phylum Mollusca, which includes snails, clams, tusk shells, and several other classes. Not all shelled ...

s. Conchology is one aspect of

malacology Malacology, from Ancient Greek μαλακός (''malakós''), meaning "soft", and λόγος (''lógos''), meaning "study", is the branch of invertebrate zoology that deals with the study of the Mollusca (molluscs or mollusks), the second-largest ...

, the study of molluscs; however, malacology is the study of molluscs as whole organisms, whereas conchology is confined to the study of their shells. It includes the study of land and freshwater mollusc shells as well as

seashell A seashell or sea shell, also known simply as a shell, is a hard, protective outer layer usually created by an animal or organism that lives in the sea. Most seashells are made by Mollusca, mollusks, such as snails, clams, and oysters ...

s and extends to the study of a

gastropod Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...

's operculum. Conchology is now sometimes seen as an archaic study, because relying on only one aspect of an organism's morphology can be misleading. However, a shell often gives at least some insight into molluscan

taxonomy image:Hierarchical clustering diagram.png, 280px, Generalized scheme of taxonomy Taxonomy is a practice and science concerned with classification or categorization. Typically, there are two parts to it: the development of an underlying scheme o ...

, and historically the shell was often the only part of exotic species that was available for study. Even in current museum collections it is common for the dry material (shells) to greatly exceed the amount of material that is preserved whole in alcohol. Conchologists mainly deal with four molluscan classes: the

s (snails),

bivalve Bivalvia () or bivalves, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class (biology), class of aquatic animal, aquatic molluscs (marine and freshwater) that have laterally compressed soft bodies enclosed b ...

s (clams), Polyplacophora (chitons) and

Scaphopoda Scaphopoda (plural scaphopods , from Ancient Greek σκᾰ́φης ''skáphē'' "boat" and πούς ''poús'' "foot"), whose members are also known as tusk shells or tooth shells, are a class of shelled marine invertebrates belonging to the ph ...

(tusk shells).

Cephalopod A cephalopod is any member of the molluscan Taxonomic rank, class Cephalopoda (Greek language, Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral symm ...

s only have small internal shells, with the exception of the Nautiloidea. Some groups, such as the sea slug

nudibranch Nudibranchs () are a group of soft-bodied marine gastropod molluscs, belonging to the order Nudibranchia, that shed their shells after their larval stage. They are noted for their often extraordinary colours and striking forms, and they have b ...

s, have lost their shells altogether, while in others it has been replaced by a

protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residue (biochemistry), residues. Proteins perform a vast array of functions within organisms, including Enzyme catalysis, catalysing metab ...

support structure.

Versus shell collecting

The terms ''shell collector'' and ''conchologist'' can be regarded as two distinct categories. Not all shell collectors are conchologists; some are primarily concerned with the

aesthetic Aesthetics (also spelled esthetics) is the branch of philosophy concerned with the nature of beauty and taste, which in a broad sense incorporates the philosophy of art.Slater, B. H.Aesthetics ''Internet Encyclopedia of Philosophy,'' , acces ...

value of shells instead of their scientific study. It is also true that not all conchologists are shell collectors; this type of research only requires access to private or institutional shell collections. There is some debate in the conchological community, with some people regarding all shell collectors (regardless of motivation) as conchologists.

History

Shell collecting, the precursor of conchology, dates back thousands of years. Archaeologists have sometimes uncovered

Stone Age The Stone Age was a broad prehistory, prehistoric period during which Rock (geology), stone was widely used to make stone tools with an edge, a point, or a percussion surface. The period lasted for roughly 3.4 million years and ended b ...

oceanic seashell necklaces in areas far from the ocean, indicating that they were traded, and shell jewellery has been found at archaeological sites around the world. During the

Renaissance The Renaissance ( , ) is a Periodization, period of history and a European cultural movement covering the 15th and 16th centuries. It marked the transition from the Middle Ages to modernity and was characterized by an effort to revive and sur ...

people began collecting natural objects of beauty for private cabinets of curiosities. Because of their attractiveness, variety, durability and ubiquity, shells frequently became a large part of such collections. Scientific interest began to develop towards the end of the 17th century, and in 1681 The Jesuit priest Filippo Bonanni published the two-volume atlas ''Ricreazione dell'occhio et della mente nell'osservazione delle chiocciole'' ("Recreation of the eye and of the mind in the observation of molluscs"), the first treatise devoted entirely to mollusc shells. In 1692 Martin Lister published '' Historia Conchyliorum'', a comprehensive conchological text with more than 1,000 engraved plates. Lister Martin conchs 1770

George Rumpf, or "Rumphius", (1627–1702) published the first genuine mollusc taxonomy. He suggested the categories "single shelled ones" (modern Polyplacophora,

limpet Limpets are a group of aquatic snails with a conical gastropod shell, shell shape (patelliform) and a strong, muscular foot. This general category of conical shell is known as "patelliform" (dish-shaped). Existing within the class Gastropoda, ...

s, and abalone), "snails or whelks" (

Gastropoda Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...

), and "two-shelled ones" (

Bivalvia Bivalvia () or bivalves, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class (biology), class of aquatic animal, aquatic molluscs (marine and freshwater) that have laterally compressed soft bodies enclosed b ...

). He did not include the tusk shells or the internal shells of the cephalopods. Many of Rumpf's terms were later adopted by

Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...

. The study of

zoology Zoology ( , ) is the scientific study of animals. Its studies include the anatomy, structure, embryology, Biological classification, classification, Ethology, habits, and distribution of all animals, both living and extinction, extinct, and ...

, including conchology, was revolutionized by Linnaeus and his system of

binomial nomenclature In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...

. Six hundred eighty three of the approximately 4,000 animal species Linnaeus described are now considered to be molluscs, although Linnaeus placed them in several different

phyla Phyla, the plural of ''phylum'', may refer to: * Phylum, a biological taxon between Kingdom and Class * by analogy, in linguistics, a large division of possibly related languages, or a major language family which is not subordinate to another Phy ...

at the time. The English word "conchology" was coined in the 1770s by the British Sephardi naturalist Emanuel Mendes da Costa, who publishe
''The Elements of Conchology: or, an Introduction to the Knowledge of Shells''
in London in 1776. Since 1700 a number of prominent conchologists have published their studies of shells. John Mawe (1764–1829) produced arguably the first conchology guidebook, ''The Voyager's Companion or Shell-Collector's Pilot'', as well as ''The Linnæan System of Conchology''. Hugh Cuming (1791–1865) is famous for his huge collection and numerous discoveries of new species.

Thomas Say Thomas Say (June 27, 1787 – October 10, 1834) was an American entomologist, conchologist, and Herpetology, herpetologist. His studies of insects and shells, numerous contributions to scientific journals, and scientific expeditions to Florida, Ge ...

wrote the fundamental work ''American Conchology, or Descriptions of the Shells of North America, Illustrated From Coloured Figures From Original Drawings, Executed from Nature'' in six volumes (1830–1834). R. Tucker Abbott was arguably the most prominent conchologist of the 20th century, authoring dozens of books and working as museum director of the Bailey-Matthews Shell Museum. His most best-known works are ''American Seashells'', ''Seashells of the World'', and ''The Kingdom of the Seashell''. John DuPont is also known for his extensive collection which he donated to the Delaware Museum of Natural History in 1984. The Japanese emperor

Hirohito , Posthumous name, posthumously honored as , was the 124th emperor of Japan according to the traditional order of succession, from 25 December 1926 until Death and state funeral of Hirohito, his death in 1989. He remains Japan's longest-reigni ...

also amassed a huge collection, and was a competent and respected amateur conchologist. In 1950, Joyce Allan, an Australian conchologist working as curator of shells at the

Australian Museum The Australian Museum, originally known as the Colonial Museum or Sydney Museum. is a heritage-listed museum at 1 William Street, Sydney, William Street, Sydney central business district, Sydney CBD, New South Wales. It is the oldest natural ...

, authored ''Australian Shells.'' This was the first book to catalogue the majority of Australian molluscs in detail and was highly regarded within the scientific community and mollusc collectors.

Museums

Many museums worldwide contain very large and scientifically important shell collections. However, in most cases these are research collections not as readily accessible to the general public as exhibits. the world's largest assemblage of mollusc shells was held by the

Smithsonian Institution The Smithsonian Institution ( ), or simply the Smithsonian, is a group of museums, Education center, education and Research institute, research centers, created by the Federal government of the United States, U.S. government "for the increase a ...

, which has c. 1 million lots representing perhaps 50,000 species. The Burke Museum of Natural History and Culture also has a large collection which was donated by Dr. Phil Nudelman in 2013. It includes about 100,000 specimens and 24,000 species, mostly from the Indo-Pacific region, the Caribbean, and the Mediterranean.

United States

* Academy of Natural Sciences,

Philadelphia Philadelphia ( ), colloquially referred to as Philly, is the List of municipalities in Pennsylvania, most populous city in the U.S. state of Pennsylvania and the List of United States cities by population, sixth-most populous city in the Unit ...

American Museum of Natural History The American Museum of Natural History (AMNH) is a natural history museum on the Upper West Side of Manhattan in New York City. Located in Theodore Roosevelt Park, across the street from Central Park, the museum complex comprises 21 interconn ...

New York City New York, often called New York City (NYC), is the most populous city in the United States, located at the southern tip of New York State on one of the world's largest natural harbors. The city comprises five boroughs, each coextensive w ...

* Bailey-Matthews Shell Museum in Sanibel Island,

Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...

: the only museum in the world dedicated entirely to shells. * Charleston Marine Life Center, Oregon Institute of Marine Biology in Charleston

Oregon Oregon ( , ) is a U.S. state, state in the Pacific Northwest region of the United States. It is a part of the Western U.S., with the Columbia River delineating much of Oregon's northern boundary with Washington (state), Washington, while t ...

* Denver Museum of Nature & Science,

Denver Denver ( ) is a List of municipalities in Colorado#Consolidated city and county, consolidated city and county, the List of capitals in the United States, capital and List of municipalities in Colorado, most populous city of the U.S. state of ...

Colorado Colorado is a U.S. state, state in the Western United States. It is one of the Mountain states, sharing the Four Corners region with Arizona, New Mexico, and Utah. It is also bordered by Wyoming to the north, Nebraska to the northeast, Kansas ...

: approximately 17,500 shell lots. * Museum of Comparative Zoology at

Cambridge Cambridge ( ) is a List of cities in the United Kingdom, city and non-metropolitan district in the county of Cambridgeshire, England. It is the county town of Cambridgeshire and is located on the River Cam, north of London. As of the 2021 Unit ...

Massachusetts Massachusetts ( ; ), officially the Commonwealth of Massachusetts, is a U.S. state, state in the New England region of the Northeastern United States. It borders the Atlantic Ocean and the Gulf of Maine to its east, Connecticut and Rhode ...

National Museum of Natural History The National Museum of Natural History (NMNH) is a natural history museum administered by the Smithsonian Institution, located on the National Mall in Washington, D.C., United States. It has free admission and is open 364 days a year. With 4.4 ...

, Washington D.C. – The Smithsonian has c. 1 million lots, the largest worldwide

Europe

Austria Austria, formally the Republic of Austria, is a landlocked country in Central Europe, lying in the Eastern Alps. It is a federation of nine Federal states of Austria, states, of which the capital Vienna is the List of largest cities in Aust ...

Vienna Vienna ( ; ; ) is the capital city, capital, List of largest cities in Austria, most populous city, and one of Federal states of Austria, nine federal states of Austria. It is Austria's primate city, with just over two million inhabitants. ...

– Naturhistorisches Museum *

Belgium Belgium, officially the Kingdom of Belgium, is a country in Northwestern Europe. Situated in a coastal lowland region known as the Low Countries, it is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeas ...

Brussels Brussels, officially the Brussels-Capital Region, (All text and all but one graphic show the English name as Brussels-Capital Region.) is a Communities, regions and language areas of Belgium#Regions, region of Belgium comprising #Municipalit ...

–

Royal Belgian Institute of Natural Sciences The Museum of Natural Sciences (, ; , ) is a Brussels museum dedicated to natural history. It is a part of the Royal Belgian Institute of Natural Sciences (; ), itself part of the institutions of the Belgian Federal Science Policy Office (BELSP ...

, one of the three largest collections *

France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...

Paris Paris () is the Capital city, capital and List of communes in France with over 20,000 inhabitants, largest city of France. With an estimated population of 2,048,472 residents in January 2025 in an area of more than , Paris is the List of ci ...

–

Muséum national d'Histoire naturelle The French National Museum of Natural History ( ; abbr. MNHN) is the national natural history museum of France and a of higher education part of Sorbonne University. The main museum, with four galleries, is located in Paris, France, within the Ja ...

900,000 lots, 5 million specimens *

Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...

Frankfurt Frankfurt am Main () is the most populous city in the States of Germany, German state of Hesse. Its 773,068 inhabitants as of 2022 make it the List of cities in Germany by population, fifth-most populous city in Germany. Located in the forela ...

– Naturmuseum Senckenberg, 700,000 lots (33,000 living taxa, 12,000 fossil taxa) **

Berlin Berlin ( ; ) is the Capital of Germany, capital and largest city of Germany, by both area and List of cities in Germany by population, population. With 3.7 million inhabitants, it has the List of cities in the European Union by population withi ...

– Berlin's Natural History Museum *

Netherlands , Terminology of the Low Countries, informally Holland, is a country in Northwestern Europe, with Caribbean Netherlands, overseas territories in the Caribbean. It is the largest of the four constituent countries of the Kingdom of the Nether ...

Leiden Leiden ( ; ; in English language, English and Archaism, archaic Dutch language, Dutch also Leyden) is a List of cities in the Netherlands by province, city and List of municipalities of the Netherlands, municipality in the Provinces of the Nethe ...

– Natural History Museum, Leiden *

Sweden Sweden, formally the Kingdom of Sweden, is a Nordic countries, Nordic country located on the Scandinavian Peninsula in Northern Europe. It borders Norway to the west and north, and Finland to the east. At , Sweden is the largest Nordic count ...

Stockholm Stockholm (; ) is the Capital city, capital and List of urban areas in Sweden by population, most populous city of Sweden, as well as the List of urban areas in the Nordic countries, largest urban area in the Nordic countries. Approximately ...

– Swedish Museum of Natural History

United Kingdom

Source: *

London London is the Capital city, capital and List of urban areas in the United Kingdom, largest city of both England and the United Kingdom, with a population of in . London metropolitan area, Its wider metropolitan area is the largest in Wester ...

–

Natural History Museum A natural history museum or museum of natural history is a scientific institution with natural history scientific collection, collections that include current and historical records of animals, plants, Fungus, fungi, ecosystems, geology, paleo ...

8 million specimens, 60,000 type specimens *

Cardiff Cardiff (; ) is the capital city, capital and List of urban areas in the United Kingdom, largest city of Wales. Cardiff had a population of in and forms a Principal areas of Wales, principal area officially known as the City and County of Ca ...

– National Museum Cardiff, second largest UK collection, over 2 million specimens *

Manchester Manchester () is a city and the metropolitan borough of Greater Manchester, England. It had an estimated population of in . Greater Manchester is the third-most populous metropolitan area in the United Kingdom, with a population of 2.92&nbs ...

–

Manchester Museum Manchester Museum is a museum displaying works of archaeology, anthropology and natural history and is owned by the University of Manchester, in England. Sited on Wilmslow Road, Oxford Road (A34 road, A34) at the heart of the university's group ...

, fourth largest UK collection; 166,000 lots. *

–

Cambridge University Museum of Zoology The University Museum of Zoology is a museum of the University of Cambridge and part of the research community of the Department of Zoology. The public is welcome and admission is free (2018). The Museum of Zoology is in the David Attenborough ...

, over 100,000 lots

Organizations

Like other scientific fields, conchologists have a number of local, national, and international organizations. There are also many organizations specializing in specific subareas. * Association Française de Conchyliologie * Belgian Society for Conchology * Club Conchylia, the German/Austrian Society for Shell Collecting * Conchological Society of Great Britain and Ireland * Conchologists of America * Conquiliologistas do Brasil * Nederlandse Malacologische Vereniging *

Unitas Malacologica Unitas may refer to: Places * 306 Unitas, a main belt asteroid * Housing complex Unitas, an apartment complex from 1931 in Bratislava, Slovakia * Johnny Unitas Stadium, Towson, Maryland, USA; a multipurpose stadium People * Johnny Unitas (1933-20 ...

Depictions of shells on stamps and coins

Shells have been featured on over 5,000

postage stamp A postage stamp is a small piece of paper issued by a post office, postal administration, or other authorized vendors to customers who pay postage (the cost involved in moving, insuring, or registering mail). Then the stamp is affixed to the f ...

s worldwide, and have been featured on many coins including the Bahamian dollar (1974), the

Cuban peso The Cuban peso ( in Spanish language, Spanish, ISO 4217 code: CUP) also known as (Spanish for 'national currency'), is the official currency of Cuba. The Cuban peso historically circulated at par with the Spanish dollar, Spanish-American silver ...

(1981), the Haitian gourde (1973), the

Nepalese rupee The Nepalese rupee (; currency sign, sign: रु; ISO 4217, code: NPR) is the official currency and legal tender of Nepal. It is also sometimes abbreviated as N₨ or Re./Rs. informally. The rupee is subdivided into 100 paisa, although coins of ...

(1989) and Philippine peso (1993).

References

Sources

National Geographic Magazine ''National Geographic'' (formerly ''The National Geographic Magazine'', sometimes branded as ''Nat Geo'') is an American monthly magazine published by National Geographic Partners. The magazine was founded in 1888 as a scholarly journal, nine ...

, March 1969, "The Magic Lure of Sea Shells", Paul A. Zahl * Seashells of the Northern Hemisphere, 1990,

Surrey Surrey () is a Ceremonial counties of England, ceremonial county in South East England. It is bordered by Greater London to the northeast, Kent to the east, East Sussex, East and West Sussex to the south, and Hampshire and Berkshire to the wes ...

, R. Tucker Abbott

External links

The Bailey-Matthews Shell Museum
in Sanibel Island
A site for amateurs
*
Beautiful Shells
' (1856) by H. G. Adams

Conchological organizations

Association Française de ConchyliologieConchologists of America

Conchological Society of Great Britain and Ireland

Club Conchylia, the German/Austrian Society for Shell Collecting

Belgian Society for Conchology

Conquiliologistas do Brasil
{{Authority control Subfields of zoology