''Combretum'', the bushwillows or combretums, make up the

type genus In biological taxonomy, the type genus is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearing type of a nominal f ...

of the family

Combretaceae The Combretaceae, often called the white mangrove family, are a family of flowering plants in the order Myrtales. The family includes about 530 species of trees, shrubs, and lianas in ca 10 genera. The family includes the leadwood tree, ''Combre ...

. The

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

comprises about 272

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

s and

shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...

s, most of which are native to

tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...

and southern

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

, about 5 to

Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...

, but there are others that are native to tropical

Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...

New Guinea New Guinea (; Hiri Motu Hiri Motu, also known as Police Motu, Pidgin Motu, or just Hiri, is a language of Papua New Guinea, which is spoken in surrounding areas of Port Moresby (Capital of Papua New Guinea). It is a simplified version of ...

and the

Bismarck Archipelago The Bismarck Archipelago (, ) is a group of islands off the northeastern coast of New Guinea in the western Pacific Ocean and is part of the Islands Region of Papua New Guinea. Its area is about 50,000 square km. History The first inhabitants o ...

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

, and tropical

America The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

. Though somewhat reminiscent of

willow Willows, also called sallows and osiers, from the genus ''Salix'', comprise around 400 speciesMabberley, D.J. 1997. The Plant Book, Cambridge University Press #2: Cambridge. of typically deciduous trees and shrubs, found primarily on moist s ...

s (''Salix'') in their habitus, they are not particularly close relatives of these.

Ecology

Bushwillow trees often are important plants in their

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

Savannah A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the Canopy (forest), canopy does not close. The open canopy allows sufficient light to rea ...

s in Africa, in particular those growing on

granitic A granitoid is a generic term for a diverse category of coarse-grained igneous rocks that consist predominantly of quartz, plagioclase, and alkali feldspar. Granitoids range from plagioclase-rich tonalites to alkali-rich syenites and from quart ...

soils, are often dominated by ''Combretum'' and its close relative ''

Terminalia Terminalia may refer to: * Terminalia (festival), a Roman festival to the god of boundaries Terminus * ''Terminalia'' (plant), a tree genus * Terminalia (insect anatomy), the terminal region of the abdomen in insects * ''Polyscias terminalia'', a ...

''. For example, ''C. apiculatum'' is a notable tree in the

Angolan mopane woodlands Angolan mopane woodlands are situated in southwestern Angola, extending into northern Namibia. This ecosystem surrounds Etosha Pan, which is considered a separate ecoregion. The mopane trees are the main type of vegetation. Location and descript ...

ecoregion in the

Kunene River The Cunene (Portuguese spelling) or Kunene (common Namibian spelling) is a river in Southern Africa. It flows from the Angola highlands south to the border with Namibia. It then flows west along the border until it reaches the Atlantic Ocean. It ...

basin in southern Africa. Other species of this genus are a major component of

Southwestern Amazonian moist forests The Southwest Amazon moist forests (NT0166) is an ecoregion located in the Upper Amazon basin. The forest is characterized by a relatively flat landscape with alluvial plains dissected by undulating hills or high terraces. The biota of the sout ...

. This genus contains several species that are

pollinated Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, ...

mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...

s other than

bat Bats are mammals of the order Chiroptera.''cheir'', "hand" and πτερόν''pteron'', "wing". With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most bi ...

s, which is quite rare indeed. But most species are more conventionally pollinated by

insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...

s or

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...

s. ''

Typhlodromus combretum ''Typhlodromus'' is a genus of predatory mites belonging to the family Phytoseiidae. Members of this genus feed largely on other mites, such as red spider mites, and several species that are popular as biological control agents to control these ...

'', a

mite Mites are small arachnids (eight-legged arthropods). Mites span two large orders of arachnids, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari, but genetic analysis does not show clear evid ...

of the family

Phytoseiidae The Phytoseiidae are a family of mites which feed on thrips and other mite species. They are often used as a biological control agent for managing mite pests. Because of their usefulness as biological control agents, interest in Phytoseiidae has ...

, was discovered on a bushwillow plant and is named after this genus. Other

herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...

s that eat ''Combretum'' foliage include the

caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...

s of the Brown Awl (''Badamia exclamationis'') which is found on '' C. albidum, C. latifolium'' and '' C. ovalifolium''; those of the Orange-tailed Awl (''Bibasis sena'') are recorded from '' C. extensum'' and ''C. latifolium''.

Use by humans

Several species are used in

African African or Africans may refer to: * Anything from or pertaining to the continent of Africa: ** People who are native to Africa, descendants of natives of Africa, or individuals who trace their ancestry to indigenous inhabitants of Africa *** Ethn ...

Indian Indian or Indians may refer to: Peoples South Asia * Indian people, people of Indian nationality, or people who have an Indian ancestor ** Non-resident Indian, a citizen of India who has temporarily emigrated to another country * South Asia ...

herbal medicine Herbal medicine (also herbalism) is the study of pharmacognosy and the use of medicinal plants, which are a basis of traditional medicine. With worldwide research into pharmacology, some herbal medicines have been translated into modern remed ...

. The class of chemical compounds known as

combretastatin Combretastatin is a dihydrostilbenoid found in '' Combretum caffrum''. As a class Combretastatins are a class of natural phenols. A variety of different natural combretastatin molecules are present in the bark of '' Combretum caffrum'', commonly ...

s were first isolated from

South African Bushwillow ''Combretum caffrum'' is the Eastern Cape South African bushwillow tree. Biochemistry In ''C. caffrum'', combretastatins A-1, A-4 and B-1 can be found.derivative In mathematics, the derivative of a function of a real variable measures the sensitivity to change of the function value (output value) with respect to a change in its argument (input value). Derivatives are a fundamental tool of calculus. F ...

, fosbretabulin disodium (combretastatin A4 phosphate), underwent preliminary study for the treatment of

anaplastic thyroid cancer Anaplastic thyroid cancer (ATC), also known as anaplastic thyroid carcinoma, is an aggressive form of thyroid cancer characterized by uncontrolled growth of cells in the thyroid gland. This form of cancer generally carries a very poor prognosis du ...

, but it was not effective enough to progress to more advanced trials. '' C. molle'' is also recorded to contain

antioxidant Antioxidants are compounds that inhibit oxidation, a chemical reaction that can produce free radicals. This can lead to polymerization and other chain reactions. They are frequently added to industrial products, such as fuels and lubricant ...

s such as

punicalagin Punicalagin is an ellagitannin, a type of phenolic compound. It is found as alpha and beta isomers in pomegranates (''Punica granatum''), ''Terminalia catappa'', ''Terminalia myriocarpa'', and in ''Combretum molle'', the velvet bushwillow, a plant ...

, which is also found in

pomegranate The pomegranate (''Punica granatum'') is a fruit-bearing deciduous shrub in the family Lythraceae, subfamily Punicoideae, that grows between tall. The pomegranate was originally described throughout the Mediterranean Basin, Mediterranean re ...

s (''Punica granatum''), a somewhat related plant. The

botanist Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...

George Don George Don (29 April 1798 – 25 February 1856) was a Scottish botanist and plant collector. Life and career George Don was born at Doo Hillock, Forfar, Angus, Scotland on 29 April 1798 to Caroline Clementina Stuart and George Don (b.1756), p ...

studied this genus extensively. The

Luvuvhu River The Levubu River or Levuvhu ( ts, Rivubye; ve, Luvuvhu) is located in the northern Limpopo province of South Africa. Some of its tributaries, such as the Mutshindudi River and Mutale River rise in the Soutpansberg Mountains. The Luvuvhu River f ...

in southern Africa was named after the river's bushwillows ('' C. erythrophyllum''), locally known as ''muvuvhu''.

Species

As of April 2021, there are 272 accepted species of ''Combretum'': *''

Combretum aculeatum ''Combretum aculeatum'' is a species of flowering plant in the bushwillow genus ''Combretum'', family Combretaceae The Combretaceae, often called the white mangrove family, are a family of flowering plants in the order Myrtales. The family inc ...

'' *''

Combretum acuminatum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum acutifolium'' *''

Combretum acutum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *''

Combretum adenogonium ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum adrianii'' *''

Combretum afzelii ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum alatum'' *''

Combretum albiflorum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *''

Combretum albopunctatum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum alfredi'' *''

Combretum andongense ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *''

Combretum andradae ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *''

Combretum angolense ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *''

Combretum angustipetalum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *''

Combretum annulatum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *''

Combretum apetalum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum aphanopetalum'' *'' Combretum apiculatum'' *''

Combretum argenteum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum argyrotrichum'' *'' Combretum assimile'' *'' Combretum atropurpureum'' *''

Combretum aureonitens ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum auriculatum'' *'' Combretum barbatum'' *'' Combretum batesii'' *'' Combretum bauchiense'' *'' Combretum baumii'' *'' Combretum bipindense'' *'' Combretum blepharopetalum'' *''

Combretum boinensis ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum bracteatum'' *'' Combretum bracteosum'' *''

Combretum brassiciforme ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum brevistylum'' *'' Combretum brunneum'' *'' Combretum butyrosum'' *'' Combretum cacoucia'' *''

Combretum caffrum ''Combretum caffrum'' is the Eastern Cape South African bushwillow tree. Biochemistry In ''C. caffrum'', combretastatins A-1, A-4 and B-1 can be found.Combretum camporum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *''

Combretum capitatum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *''

Combretum capituliflorum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum capuronii'' *''

Combretum carringtonianum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum caudatisepalum'' *''

Combretum celastroides ''Combretum celastroides'' is a species of flowering plant in the bushwillow genus ''Combretum'', family Combretaceae. It is found in Zambia, Zimbabwe, Botswana, and Namibia Namibia (, ), officially the Republic of Namibia, is a country in ...

'' *'' Combretum chinense'' *'' Combretum chionanthoides'' *''

Combretum cinereopetalum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *''

Combretum cinnabarinum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum clarense'' *''

Combretum coccineum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *''

Combretum collinum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *''

Combretum comosum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum conchipetalum'' *'' Combretum confertum'' *'' Combretum × confusum'' *''

Combretum congolanum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *''

Combretum constrictum ''Combretum constrictum'' is a flowering tree species endemic to tropical Africa, within Afrotropic ecoregions. Countries it is native to include: Kenya; Tanzania, Somalia, Mozambique, and Nigeria Nigeria ( ), , ig, Naìjíríyà, yo, ...

'' *'' Combretum contractum'' *'' Combretum coriifolium'' *'' Combretum coursianum'' *'' Combretum cuspidatum'' *'' Combretum decandrum'' *''

Combretum decaryi ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum deciduum'' *'' Combretum demeusei'' *'' Combretum discolor'' *''

Combretum dolichopodum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum duarteanum'' *''

Combretum echirense ''Combretum echirense'' is a species of flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('conta ...

'' *''

Combretum edwardsii ''Combretum edwardsii'', the Natal combretum or forest climbing bushwillow, is an uncommon forest plant endemic to the mistbelt region of eastern South Africa. It has a climbing habit and the stems may often lie prostrate on the forest floor or o ...

'' *''

Combretum elaeagnoides ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum engleri'' *''

Combretum eriogynum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...

'' *''

Combretum erosum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *''

Combretum erythrophloeum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *''

Combretum erythrophyllum ''Combretum erythrophyllum'', commonly known as the river bushwillow, is a medium to large-sized, spreading tree found in bush along river banks in southern Africa. It is planted as a shade and ornamental tree in South Africa and the United State ...

'' *'' Combretum esteriense'' *''

Combretum evisceratum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum exalatum'' *'' Combretum exannulatum'' *''

Combretum excelsum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum exellii'' *'' Combretum falcatum'' *'' Combretum farinosum'' *'' Combretum flammeum'' *'' Combretum foliatum'' *''

Combretum frangulifolium ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *''

Combretum fruticosum ''Combretum fruticosum'', known as orange flame vine or chameleon vine, is a species of bushwillow that occurs from Mexico to northern Argentina. Description The orange flame vine is a liana that reaches up to in height without a support a ...

'' *''

Combretum fulvum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...

'' *''

Combretum fuscum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum gabonense'' *'' Combretum germainii'' *''

Combretum ghesquierei ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum gillettianum'' *''

Combretum glabrum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum glaucocarpum'' *''

Combretum glutinosum ''Combretum glutinosum'' is a shrub species of the genus ''Combretum'', found in the Sahel belt in parts of Senegal, Burkina Faso, Ghana, Mali, the Gambia, Niger, Nigeria and Cameroon, across to parts of Sudan. It is known as dooki in Pulaar, Ka ...

'' *''

Combretum goetzei ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...

'' *'' Combretum goldieanum'' *'' Combretum goossensii'' *'' Combretum gordonii'' *'' Combretum gossweileri'' *''

Combretum gracile ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum graciliflorum'' *'' Combretum grandidieri'' *'' Combretum grandiflorum'' *'' Combretum griffithii'' *'' Combretum harmsianum'' *'' Combretum harrisii'' *'' Combretum hartmannianum'' *'' Combretum haullevilleanum'' *''

Combretum hensii ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *''

Combretum hereroense ''Combretum hereroense'', commonly known as the russet bushwillow and the mouse-eared combretum, is a deciduous shrub or small tree that is found from eastern Africa to northern South Africa. Over its extensive range it is variable with respect t ...

'' *'' Combretum hilarianum'' *''

Combretum holstii ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *''

Combretum homalioides ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...

'' *''

Combretum igneiflorum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum illairii'' *''

Combretum imberbe ''Combretum imberbe'' (leadwood, af, hardekool, st, mohwelere-tšhipi, ts, motswiri/mondzo, zu, impondondlovu) is a characteristic and often impressive bushwillow species of the southern Afrotropics. The medium to large tree has a sparse, ...

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Combretum indicum ''Combretum indicum'', also known as the Rangoon creeper, is a vine with red flower clusters which is native to tropical Asia. Description The Rangoon creeper is a ligneous vine that can reach from 2.5 meters to up to 8 meters. The leaves are e ...

'' *'' Combretum inflatum'' *'' Combretum ivanii'' *'' Combretum kasaiense'' *''

Combretum kirkii ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

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Combretum klossii ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum klotzschii'' *'' Combretum kostermansii'' *'' Combretum kraussii'' *''

Combretum lanceolatum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum lanuginosum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum latialatum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...

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Combretum latifolium ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...

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Combretum laxum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum lecardii'' *''

Combretum leprosum ''Combretum leprosum'', the mofumbo in Portuguese and the asucaró in eastern Bolivian Spanish, is a plant species in the genus ''Combretum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The ...

'' *'' Combretum lindense'' *''

Combretum lisowskii ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

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Combretum littoreum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

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Combretum llewelynii ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...

'' *'' Combretum lokele'' *'' Combretum longicollum'' *''

Combretum longipilosum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum longispicatum'' *''

Combretum longistipitatum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum louisii'' *'' Combretum lukafuensis'' *''

Combretum luxenii ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

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Combretum macrocalyx ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum malabaricum'' *'' Combretum mannii'' *'' Combretum marginatum'' *''

Combretum marquesii ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum mellifluum'' *''

Combretum meridionalis ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum micranthum ''Combretum micranthum'' is a species of flowering plant in the family Combretaceae. It is a shrub known as kinkeliba in Benin, Burkina Faso, Senegal, Mali and the Gambia across multiple regional dialects, where it is often found in tiger bush ...

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Combretum microphyllum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum mkuzense'' *''

Combretum moggii ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum molle ''Combretum molle'', the velvet bushwillow, is a medium to large tree species in the genus ''Combretum'' found in western, eastern and southern Africa. The larvae of '' Parosmodes morantii'' and '' Acalyptris molleivora'' feed on ''C. molle''. ...

'' *'' Combretum monetaria'' *''

Combretum mooreanum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

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Combretum mortehanii ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum mossambicense ''Combretum mossambicense'' is a climbing shrub occurring in Eastern Zambia, Zimbabwe and Mozambique Mozambique (), officially the Republic of Mozambique ( pt, Moçambique or , ; ny, Mozambiki; sw, Msumbiji; ts, Muzambhiki), is a count ...

'' *'' Combretum mucronatum'' *'' Combretum multinervium'' *'' Combretum mussaendiflorum'' *''

Combretum mweroense ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...

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Combretum nanum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...

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Combretum ndjoleense ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

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Combretum nigrescens ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum nigricans'' *'' Combretum nioroense'' *''

Combretum niphophilum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...

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Combretum nusbaumeri ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...

'' *'' Combretum oatesii'' *'' Combretum obovatum'' *'' Combretum obscurum'' *'' Combretum octagonum'' *''

Combretum olivaceum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

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Combretum oudenhovenii ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum ovalifolium ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum oxygonium ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

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Combretum oxystachyum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum oyemense ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...

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Combretum padoides ''Combretum padoides'', the thicket bushwillow, occurs in the lowlands of tropical and south-eastern Africa. They grow in a range of habitats from muddy riverbanks to dry rocky hillsides. The mostly opposite oval leaves are carried on long slend ...

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Combretum paniculatum ''Combretum paniculatum'', the burning bush or forest flame-creeper, is a plant species in the genus ''Combretum'' found in Africa. The fruit is a samara, i.e. a winged seed. Chemistry The ethyl acetate Ethyl acetate ( systematically ethyl ...

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Combretum paradoxum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum paraguariense'' *''

Combretum patelliforme ''Combretum celastroides'' is a species of flowering plant in the bushwillow genus ''Combretum'', family Combretaceae. It is found in Zambia, Zimbabwe, Botswana, and Namibia, and is also known in English as Jesse-bush bushwillow, Trailing bushwil ...

'' *'' Combretum paucinervium'' *'' Combretum pavonii'' *''

Combretum pecoense ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum pellegrinianum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum pentagonum'' *'' Combretum petrophilum'' *''

Combretum phaeocarpum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum pilosum'' *'' Combretum pisoniiflorum'' *'' Combretum pisonioides'' *'' Combretum platypterum'' *'' Combretum polyanthum'' *'' Combretum polystictum'' *''

Combretum procursum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum psidioides'' *''

Combretum punctatum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum purpureiflorum'' *''

Combretum pyramidatum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

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Combretum pyrifolium ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...

'' *'' Combretum quadrangulare'' *'' Combretum quadratum'' *'' Combretum rabiense'' *''

Combretum racemosum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

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Combretum razianum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum recurvatum'' *''

Combretum relictum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum rhodanthum'' *'' Combretum riggenbachianum'' *'' Combretum robustum'' *'' Combretum robynsii'' *''

Combretum rochetianum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...

'' *'' Combretum rohrii'' *'' Combretum rotundifolium'' *'' Combretum rovirosae'' *'' Combretum roxburghii'' *'' Combretum rupestre'' *''

Combretum rupicola ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum sanjappae'' *''

Combretum scandens ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum schumannii ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

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Combretum sericeum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum shivannae ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum sordidum'' *''

Combretum sphaeroides ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum spinosum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum stenopterum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

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Combretum stocksii ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum struempellianum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...

'' *'' Combretum stylesii'' *'' Combretum subglabratum'' *''

Combretum subumbellatum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum sundaicum'' *''

Combretum sylvicola ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum tarquense ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum tenuipetiolatum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum tessmannii'' *'' Combretum tetragonocarpum'' *'' Combretum tetralophoides'' *''

Combretum tetralophum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

'' *'' Combretum teuschii'' *''

Combretum tibatiense ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum tomentosum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum towaense ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum trichophyllum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum trifoliatum ''Combretum trifoliatum'' is a vine (rarely a shrub) of the family Combretaceae. It is found from Myanmar across Southeast Asia and Wallacea to New Guinea and Australia. It grows in wet places, including where it can be submerged for four months ...

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Combretum ulei ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...

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Combretum umbricola ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but th ...

'' *'' Combretum vendae'' *'' Combretum vernicosum'' *''

Combretum villosum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

'' *'' Combretum violaceum'' *'' Combretum virgatum'' *'' Combretum viscosum'' *'' Combretum wallichii'' *'' Combretum wattii'' *''

Combretum wilksii ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

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Combretum winitii ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...

'' *'' Combretum xanthothyrsum'' *'' Combretum youngii'' *''

Combretum zenkeri ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ther ...

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Combretum zeyheri ''Combretum zeyheri'', the large-fruited bushwillow or Zeyher's bushwillow, is a species of flowering plant in the family Combretaceae, usually found growing on acidic or sandy soils in tropical African savannas. A small to medium-sized tree, its ...

References

External links

{{Taxonbar, from=Q1777188 Medicinal plants Myrtales genera