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''Pachydactylus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
insectivorous A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant that eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores were ...
gecko Geckos are small, mostly carnivorous lizards that have a wide distribution, found on every continent except Antarctica. Belonging to the infraorder Gekkota, geckos are found in warm climates throughout the world. They range from . Geckos ar ...
s,
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
s in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Gekkonidae. The genus is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
, and member
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
are commonly known as thick-toed geckos. The genus also displays rich
speciation Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution within ...
, having 57 distinct species identified when compared to other closely related gecko genera like ''
Rhoptropus ''Rhoptropus'' is a genus of geckos endemic to Southern Africa, better known as Namib day geckos. Classification of genus ''Rhoptropus'' The following species are recognized: www.reptile-database.org. *'' Rhoptropus afer'' W. Peters, 1869 — ...
,'' most of which have emerged since 35 Ma. It has been suggested that the reason for this rich speciation not from
adaptive radiation In evolutionary biology, adaptive radiation is a process in which organisms diversify rapidly from an ancestral species into a multitude of new forms, particularly when a change in the environment makes new resources available, alters biotic int ...
nor
nonadaptive radiation Nonadaptive radiations are a subset of evolutionary radiations (or species flocks) that are characterized by diversification that is not driven by resource partitioning. The species that are a part of a nonadaptive radiation will tend to have very s ...
, but that the genus represents a clade somewhere between the two drivers of speciation. ''P. bibronii'' geckos have been used by NASA as animal models for experimentation.


Description

The genus ''Pachydactylus'' is characterized by dilated toe tips, usually with undivided scansors. Body scales are small, granular and non-overlapping, with scattered, large keeled tubercles. Coloration of Pachydactylus species varies, but is generally drab in color. Presence of adhesive toe pads varies by species and habitat, with rock dwelling species of ''Pachydactylus'' retaining adhesive pads, but unambiguous independent loss of toe pads in sand dwelling and burrowing species like ''P. rangei''. Body size in ''Pachydactylus'' varies across the 57 species, ranging from 35 to 115mm Snout-Vent Length (SVL) with the ancestral condition of a larger body size with adhesive toe pads to suit a generalist habitat.


Behavior

All observed
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of ''Pachydactylus'' are strictly
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
.


Habitat

''Pachydactylus'' species live in a diverse range of
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s across Southern Africa. Habitat varies by species, with some species preferring generalist habitats, human dwellings, rock-dwellings, and sand dwellings. Habitat preference typically varies by body size and retention of toe pads, which varies across the genus. The body size of ''Pachydactylus'' geckos has been shown to correlate with their habitat range.


Diet

Lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
s of the genus ''Pachydactylus'' feed mainly on
arthropods Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
, but have been observed eating small vertebrates.


Geographic range

The geographic range of the genus ''Pachydactylus'' is centered on
Southern Africa Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number of ...
, with some species reaching
East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historical ...
, the northernmost limit of their distribution. In South Africa's rugged
Richtersveld The Richtersveld is a desert landscape characterised by rugged kloofs and high mountains, situated in the north-western corner of South Africa’s Northern Cape province. It is full of changing scenery from flat, sandy, coastal plains, to crag ...
region, ''Pachydactylus'' geckos comprised 13 of 18 all gecko species surveyed.


Species

There are 57
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
that are recognized as being valid:"''Pachydactylus'' ". The Reptile Database. www.reptile-database.org. *'' Pachydactylus acuminatus'' *''
Pachydactylus affinis ''Pachydactylus affinis'', also known as Transvaal gecko or Transvaal thick-toed gecko, is a species of lizard in the family Gekkonidae. It is found in South Africa and Zimbabwe Zimbabwe (), officially the Republic of Zimbabwe, is a land ...
'' –
Transvaal Transvaal is a historical geographic term associated with land north of (''i.e.'', beyond) the Vaal River in South Africa. A number of states and administrative divisions have carried the name Transvaal. * South African Republic (1856–1902; af, ...
thick-toed gecko *'' Pachydactylus amoenus'' *'' Pachydactylus angolensis'' - Angola large-scaled gecko, Angolan thick-toed gecko *'' Pachydactylus atorquatus'' *'' Pachydactylus austeni'' – Austen's thick-toed gecko *''
Pachydactylus barnardi ''Pachydactylus barnardi'', also known commonly as Barnard's rough gecko or Barnard's thick-toed gecko, is a species of lizard in the family Gekkonidae. The species is indigenous to Southern Africa. Etymology The specific name, ''barnardi'', ...
'' – Barnard's thick-toed gecko *'' Pachydactylus bicolor'' – velvety thick-toed gecko *'' Pachydactylus boehmei'' *''
Pachydactylus capensis ''Pachydactylus capensis'', also known as Cape gecko or Cape thick-toed gecko, is a species of lizard in the family Gekkonidae Gekkonidae (the common geckos) is the largest family of geckos, containing over 950 described species in 64 genera. ...
'' – Cape thick-toed gecko *''
Pachydactylus caraculicus The Angola banded thick-toed gecko (''Pachydactylus caraculicus'') is a species of lizard in the family Gekkonidae. It is found in southwestern Angola and northern Namibia Namibia (, ), officially the Republic of Namibia, is a country in ...
'' –
Angola , national_anthem = " Angola Avante"() , image_map = , map_caption = , capital = Luanda , religion = , religion_year = 2020 , religion_ref = , coordina ...
banded thick-toed gecko *'' Pachydactylus carinatus'' *'' Pachydactylus etultra'' *'' Pachydactylus fasciatus'' – banded thick-toed gecko *'' Pachydactylus formosus'' – Smith's thick-toed gecko *'' Pachydactylus gaiasensis'' – Brandberg thick-toed gecko *'' Pachydactylus geitje'' – ocellated thick-toed gecko *'' Pachydactylus griffini'' *'' Pachydactylus haackei'' – Haacke's thick-toed gecko *'' Pachydactylus katanganus'' *'' Pachydactylus kladaroderma'' – thin-skinned thick-toed gecko *''
Pachydactylus kobosensis ''Pachydactylus kobosensis'' is a species of lizard in the family Gekkonidae. It is endemic to Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares l ...
'' *''
Pachydactylus kochii The Cape Cross thick-toed gecko (''Pachydactylus kochii''), also known commonly as Koch's gecko and Koch's thick-toed gecko, is a species of gecko, a lizard in the family Gekkonidae. The species is endemic to southern Africa. Etymology The spe ...
'' *''
Pachydactylus labialis ''Pachydactylus labialis'', commonly known as the Calvinia thick-toed gecko, Western Cape gecko, or Western Cape thick-toed gecko, is a gecko species endemic to the Western and Northern Cape in South Africa, often found taking shelter under stone ...
'' –
Calvinia Calvinia is a regional town in the Northern Cape province of South Africa named after the French people, French religious reformer Jean Calvin. The town falls under the Hantam Local Municipality which forms part of the Namakwa District Municipalit ...
thick-toed gecko *'' Pachydactylus latirostris'' – quartz gecko *'' Pachydactylus macrolepis'' – large-scaled banded gecko *'' Pachydactylus maculatus'' – spotted thick-toed gecko *'' Pachydactylus maraisi'' *'' Pachydactylus mariquensis'' – Marico thick-toed gecko *'' Pachydactylus mclachlani'' *'' Pachydactylus monicae'' *'' Pachydactylus montanus'' *'' Pachydactylus namaquensis'' – Namaqua thick-toed gecko *'' Pachydactylus oculatus'' – golden spotted thick-toed gecko *'' Pachydactylus oreophilus'' –
Kaokoveld The Kaokoveld Desert is a coastal desert of northern Namibia and southern Angola. Setting The Kaokoveld Desert occupies a coastal strip covering , from 13° to 21°S and is bounded by the Atlantic Ocean to the west, the Namibian savanna woodla ...
thick-toed gecko *''
Pachydactylus oshaughnessyi ''Pachydactylus oshaughnessyi'' is a species of lizard in the family Gekkonidae. The species is endemic to southern Africa. Etymology The specific name, ''oshaughnessyi'', is in honor of Arthur O'Shaughnessy, who was a British herpetologist an ...
'' *'' Pachydactylus otaviensis'' *'' Pachydactylus parascutatus'' *'' Pachydactylus punctatus'' – speckled thick-toed gecko *'' Pachydactylus purcelli'' *'' Pachydactylus rangei'' – Namib sand gecko *'' Pachydactylus reconditus'' *''
Pachydactylus robertsi ''Pachydactylus robertsi'', commonly known as the large-scaled gecko, shielded thick-toed gecko, or ''Rauhschuppen-Dickfingergecko'' in German, is an African species of gecko. Geographic range ''P. robertsi'' is found near Keetmanshoop and Kara ...
'' *'' Pachydactylus rugosus'' – rough thick-toed gecko *'' Pachydactylus sansteynae'' – San Steyn's thick-toed gecko *'' Pachydactylus scherzi'' – Sherz's thick-toed gecko *''
Pachydactylus scutatus ''Pachydactylus scutatus'', also known as the large-scaled gecko or shielded thick-toed gecko, is a species of lizard in the family Gekkonidae. It is found in Namibia, South Africa, and Angola , national_anthem = " Angola Avante"( ...
'' – large-scaled thick-toed gecko *'' Pachydactylus serval'' – western spotted thick-toed gecko *'' Pachydactylus tigrinus'' – tiger thick-toed gecko *'' Pachydactylus tsodiloensis'' – Tsodilo thick-toed gecko *''
Pachydactylus vansoni ''Pachydactylus vansoni'', Common name, commonly known as Van Son's gecko or Van Son's thick-toed gecko, is a species of lizard in the Family (biology), family Gekkonidae. The species is Endemism, endemic to Southern Africa. Etymology The Specif ...
'' – Van Son's thick-toed gecko *'' Pachydactylus vanzyli'' *'' Pachydactylus visseri'' *'' Pachydactylus wahlbergii'' *'' Pachydactylus waterbergensis'' *''
Pachydactylus weberi Weber's thick-toed gecko (''Pachydactylus weberi'') is a species of lizard in the family Gekkonidae. The species is native to southern Africa. Etymology The specific name, ''weberi'', is in honor of German-Dutch zoologist Max Wilhelm Carl Web ...
'' – Weber's thick-toed gecko *'' Pachydactylus werneri'' ''
Nota bene (, or ; plural form ) is a Latin phrase meaning "note well". It is often abbreviated as NB, n.b., or with the ligature and first appeared in English writing . In Modern English, it is used, particularly in legal papers, to draw the atten ...
'': A
binomial authority In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
in parentheses indicates that the species was originally described in a genus other than ''Pachydactylus''.


References


Further reading

* * * *


External links

* {{Taxonbar, from=Q2482361 Geckos of Africa Lizard genera Taxa named by Arend Friedrich August Wiegmann