Cochliopid
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Cochliopidae is a family of small
freshwater snail Freshwater snails are gastropod mollusks which live in fresh water. There are many different families. They are found throughout the world in various habitats, ranging from ephemeral pools to the largest lakes, and from small seeps and springs ...
s with gills and an operculum, aquatic
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusks. Paludestrina d'Orbigny, 1840 is an archaic synonym, and has been placed on the Official Index of Rejected and Invalid Names by
ICZN The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its publisher, the I ...
Opinion 2202.MolluscaBase eds. (2021). MolluscaBase. Cochliopidae Tryon, 1866. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=411706 on 2021-05-13 This family is in the superfamily Truncatelloidea and in the order Littorinimorpha (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).


Description

Cochliopidae snails are characterized by sharp, elongated spire-like structures, and can be found in rivers or brackish water.


2005 taxonomy

The family Cochliopidae consists of 3 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005): * Cochliopinae Tryon, 1866 - synonyms: Mexithaumatinae D. W. Taylor, 1966, Paludiscalinae D. W. Taylor, 1966 * Littoridininae
Thiele Thiele is a German-language surname. Geographical distribution As of 2014, 78.0% of all known bearers of the surname ''Thiele'' were residents of Germany, 10.9% of the United States, 2.3% of Australia, 2.0% of Brazil, 1.0% of Canada and 1.0% of Sou ...
, 1928
* Semisalsinae Guiusti & Pezzoli, 1980 - synonym: Heleobiini Bernasconi, 1991


Genera

Liu et al. (2001) have recognized 34 genera with more than 260 species within the subfamily Cochliopinae. Strong et al. (2008)Strong E. E., Gargominy O., Ponder W. F. & Bouchet P. (2008). "Global Diversity of Gastropods (Gastropoda; Mollusca) in Freshwater". ''
Hydrobiologia ''Hydrobiologia'', ''The International Journal of Aquatic Sciences'', is a peer-reviewed scientific journal publishing 21 issues per year, for a total of well over 4000 pages per year. ''Hydrobiologia'' publishes original research, reviews and opi ...
'' 595: 149-166. .
have recognized 246 freshwater species within Cochliopidae. Genera within the family Cochliopidae include: subfamily Cochliopinae *''
Aphaostracon ''Aphaostracon'' is a genus of very small or minute freshwater and brackish water snails that have an operculum, aquatic operculate gastropod mollusks in the family Cochliopidae.Bouchet, P. (2014). Aphaostracon F. G. Thompson, 1968. Accessed th ...
'' F. G. Thompson, 1968 *''
Coahuilix ''Coahuilix'' is a genus of small freshwater snails that have an operculum, aquatic gastropod mollusks in the family Cochliopidae.Bouchet, P. (2014). Coahuilix D. W. Taylor, 1966. Accessed through: World Register of Marine Species at http://www ...
'' Taylor, 1966 *''
Cochliopa ''Cochliopa'' is a genus of freshwater snails, specifically cave snails, aquatic animal, aquatic gastropod mollusks in the family Cochliopidae. ''Cochliopa'' is the type genus of the family Cochliopidae. This genus is not to be confused with th ...
'' Stimpson, 1865 - type genus of the family Cochliopidae *''
Cochliopina ''Cochliopina'' is a genus of very small freshwater snails that have an operculum, aquatic gastropod mollusks in the family Hydrobiidae, the mud snails. Species Species within the genus ''Cochliopina'' include:: *''Cochliopina compacta'' Pilsb ...
'' Morrison, 1946 subfamily Littoridininae *''Antrobia'' Hubricht, 1971 - with the only species '' Antrobia culveri'' Hubricht, 1971Kabat A. R. & Hershler R. (1993). "The prosobranch snail family Hydrobiidae (Gastropoda: Rissooidea): review of classification and supraspecific taxa". '' Smithsonian Contributions to Zoology'' 547: 1-94
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- Tumbling Creek cavesnail *'' Littoridina'' Souleyet, 1852 *''
Pseudotryonia ''Pseudotryonia'' is a genus of gastropods belonging to the family Cochliopidae. The species of this genus are found in Northern America. Species: *''Pseudotryonia adamantina'' *''Pseudotryonia alamosae'' *''Pseudotryonia brevissima'' *''Pse ...
'' Hershler, 2001 **'' Pseudotryonia adamantina'' **'' Pseudotryonia alamosae'' **'' Pseudotryonia brevissima'' **'' Pseudotryonia grahamae'' **'' Pseudotryonia mica'' Hershler, Liu & Landye, 2011Hershler R., Liu H.-P. & Landye J. J. (2011). "Two new genera and four new species of freshwater cochliopid gastropods (Rissooidea) from northeastern Mexico". '' Journal of Molluscan Studies'' 77(1): 8-23. . **'' Pseudotryonia pasajae'' Hershler, Liu & Landye, 2011 subfamily Semisalsinae - there are three genera in the subfamily SemisalsinaeKroll O., Hershler R., Albrecht C., Terrazas E. M., Apaza R., Fuentealba C., Wolff C. & Wilke T. (2012). "The endemic gastropod fauna of Lake Titicaca: correlation between molecular evolution and hydrographic history". '' Ecology and Evolution'' 2(7): 1517-1530. . *''
Heleobia ''Heleobia'' is a genus of small freshwater and brackish water snails with a gill and an operculum, aquatic gastropod mollusks in the family Cochliopidae and the superfamily Truncatelloidea.Bouchet, P.; Gofas, S. (2015). Heleobia. Accessed thro ...
'' Stimpson, 1865 *''
Heleobops ''Heleobops'' is a genus of very small aquatic snails, operculate gastropod mollusks in the family Cochliopidae. ''Heleobops'' is one of three genera (together with ''Heleobia'' and '' Semisalsa'') within the subfamily Semisalsinae. Species Sp ...
'' Thompson, 1968 *''
Semisalsa ''Eupaludestrina'' is a genus of gastropods belonging to the family Cochliopidae. The species of this genus are found in Europe. Species: *'' Eupaludestrina achaja'' *'' Eupaludestrina acilacustris'' *''Eupaludestrina aponensis'' *'' Eupalu ...
'' Radoman, 1974 subfamily ? (either Cochliopinae or Littoridininae) *'' Aroapyrgus'' H. B. Baker, 1931 *'' Balconorbis'' Hershler & Longley, 1986 *† '' Carinulorbis'' Yen, 1949 *'' Chorrobius'' Hershler, Liu & Landye, 2011 *'' Dyris'' Conrad, 1871 - it has extant species and also 26 species in Miocene Pebas Formation *'' Emmericiella''
Pilsbry Henry Augustus Pilsbry (7 December 1862 – 26 October 1957) was an American biologist, malacologist and carcinologist, among other areas of study. He was a dominant presence in many fields of invertebrate taxonomy for the better part of a centu ...
, 1909
*'' Eremopyprgus'' Hershler, 1999Hershler R., Liu H.-P. & Landye J. J. (2002). "A new species of ''Eremopyprgus'' (Hydrobiidae: Cochliopinae) from the Chihuahuan desert, Mexico: Phylogentic relationships and biogeography". '' Journal of Molluscan Studies'' 68: 7-13
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Hershler R. (1999). "A systematic review of the hydrobiid snails (Gastropoda: Rissoidea) of the Great Basin, western United States. Part II. Genera ''Colligyrus'', ''Fluminicola'', ''Pristinicola'', and ''Tryonia''". ''The Veliger'' 42(4): 306-337
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*† '' Feliconcha'' Wesselingh, Anderson & Kadolsky, 2006 - two species from Miocene of the Pebas FormationWesselingh F. P., Anderson L. C. & Kadolsky D. (2006). "Molluscs from the Miocene Pebas Formation of Peruvian and Colombian Amazonia". '' Scripta Geologica'' 1333: 19-290
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**'' Feliconcha feliconcha'' Wesselingh, Anderson & Kadolsky, 2006 **'' Feliconcha reticulata'' Wesselingh, Anderson & Kadolsky, 2006 *(probably extant) '' Glabertryonia'' Wesselingh, Anderson & Kadolsky, 2006 - three species **† '' Glabertryonia glabra'' Wesselingh, Anderson & Kadolsky, 2006 - from Miocene of the Pebas Formation **† ''Glabertryonia'' sp. 1 - from Pliocene of the Las Piedras Formation **(probably extant) ''Glabertryonia'' sp. 2 - from Holocene of Surinam, probably extant *'' Juturnia'' Hershler, Liu & Stockwell, 2002 *'' Lithococcus''
Pilsbry Henry Augustus Pilsbry (7 December 1862 – 26 October 1957) was an American biologist, malacologist and carcinologist, among other areas of study. He was a dominant presence in many fields of invertebrate taxonomy for the better part of a centu ...
, 1911
*'' Mesobia'' F. G. Thompson & Hershler, 1991 *''
Mexipyrgus ''Mexipyrgus'' is a genus of very small freshwater snails, aquatic gastropod mollusks in the family Cochliopidae.Bouchet, P. (2014). Mexipyrgus D. W. Taylor, 1966. Accessed through: World Register of Marine Species at http://www.marinespecies ...
'' Taylor, 1966 *''
Minckleyella ''Minckleyella'' is a monotypic genus of gastropods belonging to the family Cochliopidae. The only species is ''Minckleyella balnearis''. This freshwater snail genus is named in honor of Wendell L. Minckley Wendell Lee Minckley (November 13, ...
'' Hershler, Liu & Landye, 2011 - with the only species '' Minckleyella balnearis'' Hershler, Liu & Landye, 2011 * '' Onobops'' Thompson, 1968 * '' Pyrgophorus'' Ancey, 1888 * '' Sioliella'' Haas, 1949 * ''
Spurwinkia ''Spurwinkia'' is a genus of very small aquatic snails, operculate gastropod mollusks in the family Cochliopidae.Bouchet, P. (2014). Spurwinkia Davis, Mazurkiewicz & Mandracchia, 1982. Accessed through: World Register of Marine Species at ...
'' Davis, Mazurkiewicz & Mandracchia, 1982 * ''
Texadina ''Texadina'' is a genus of very small aquatic snails, operculate gastropod mollusks in the family Cochliopidae.Bouchet, P. (2015). Texadina Abbott & Ladd, 1951. Accessed through: World Register of Marine Species at http://www.marinespecies ...
'' Abbott & Ladd, 1951 * '' Thalassobia'' Bourguignat in Mabille, 1877


Cladogram

A cladogram based on sequences of mitochondrial cytochrome-c oxidase I (COI) genes showing phylogenic relations of species within Cochliopidae:Liu H.-P., Hershler R. & Thompson F. G. (2001). "Phylogenetic Relationships of the Cochliopinae (Rissooidea: Hydrobiidae): An Enigmatic Group of Aquatic Gastropods". '' Molecular Phylogenetics and Evolution'' 21(1): 17-25. .


References


Further reading

* Hershler R., Davis C. L., Kitting C. L & Liu H.-P. (2007). "Discovery of introduced and cryptogenic cochliopid gastropods in the San Francisco Estuary, California". '' Journal of Molluscan Studies'' 73(4): 323-332
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* Hershler R. & Thompson F. G. (1992). "A Review of the Aquatic Gastropod Subfamily Cochliopinae (Prosobranchia: Hydrobiidae)". '' Malacological Review'', Supplement 5: 1-140. pages 16–18.
Wilke T., Haase M., Hershler R., Liu H.-P., Misof B. & Ponder W. (2013) ''Pushing short DNA fragments to the limit: Phylogenetic relationships of ‘hydrobioid’ gastropods (Caenogastropoda: Rissooidea)''. Molecular Phylogenetics and Evolution, 66: 715-736


External links


Cochliopidae on WoRMS
{{Taxonbar, from=Q969214