''Cobitis ohridana'' is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...

in the

true loach Cobitidae, also known as the True loaches, is a family of Old World freshwater fish. They occur throughout Eurasia and in Morocco, and inhabit riverine ecosystems. Today, most "loaches" are placed in other families (see below). The family include ...

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

(Cobitidae). It is only found in

Albania Albania ( ; sq, Shqipëri or ), or , also or . officially the Republic of Albania ( sq, Republika e Shqipërisë), is a country in Southeastern Europe. It is located on the Adriatic and Ionian Seas within the Mediterranean Sea and shares ...

Montenegro ) , image_map = Europe-Montenegro.svg , map_caption = , image_map2 = , capital = Podgorica , coordinates = , largest_city = capital , official_languages = M ...

and

North Macedonia North Macedonia, ; sq, Maqedonia e Veriut, (Macedonia before February 2019), officially the Republic of North Macedonia,, is a country in Southeast Europe. It gained independence in 1991 as one of the successor states of Socialist Feder ...

.Crivelli (2005) This fish was long believed to be part of the widespread

Spined Loach The spined loach (''Cobitis taenia'') is a common freshwater fish in Europe. It is sometimes known as spotted weather loach, not to be confused with the "typical" weather loaches of the genus ''Misgurnus''. This is the type species of the spiny l ...

(''C. taenia'').

Cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...

analysis of

DNA sequence DNA sequencing is the process of determining the nucleic acid sequence – the order of nucleotides in DNA. It includes any method or technology that is used to determine the order of the four bases: adenine, guanine, cytosine, and thymine. Th ...

data (

nDNA Nuclear DNA (nDNA), or nuclear deoxyribonucleic acid, is the DNA contained within each cell nucleus of a eukaryotic organism. It encodes for the majority of the genome in eukaryotes, with mitochondrial DNA and plastid DNA coding for the rest. I ...

RAG-1 The recombination-activating genes (RAGs) encode parts of a protein complex that plays important roles in the rearrangement and recombination of the genes encoding immunoglobulin and T cell receptor molecules. There are two recombination-activ ...

and S7

ribosomal protein A ribosomal protein (r-protein or rProtein) is any of the proteins that, in conjunction with rRNA, make up the ribosomal subunits involved in the cellular process of translation. ''E. coli'', other bacteria and Archaea have a 30S small subunit an ...

intron An intron is any nucleotide sequence within a gene that is not expressed or operative in the final RNA product. The word ''intron'' is derived from the term ''intragenic region'', i.e. a region inside a gene."The notion of the cistron .e., gene. ...

1, and

mtDNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA ...

cytochrome ''b'') confirms that it is properly treated as full species. Its closest living relative might be '' C. zanandreai'' of

Lake Fondi Lake Fondi ( it, Lago di Fondi, la, Lacus Fundanus, Lacus Amyclanus) is a brackish lake about to the southeast of Rome in the Province of Latina, Lazio, Italy, in the region called Sud or "South" Pontino, the western end of which is the Piana di ...

and its surroundings in west-central

Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...

. In that case these species separated probably around 6 million years ago in the

Late Miocene The Late Miocene (also known as Upper Miocene) is a sub-epoch of the Miocene epoch (geology), Epoch made up of two faunal stage, stages. The Tortonian and Messinian stages comprise the Late Miocene sub-epoch, which lasted from 11.63 Ma (million ye ...

, perhaps during the early

Messinian salinity crisis The Messinian salinity crisis (MSC), also referred to as the Messinian event, and in its latest stage as the Lago Mare event, was a geological event during which the Mediterranean Sea went into a cycle of partial or nearly complete desiccation (dr ...

. Alternatively, these two might simply be independently derived from a common ancestor in the course of these events but probably somewhat earlier, and appear similar on the molecular level due to

convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...

.Perdices ''et al.'' (2008) Its natural

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

s are

Lake Ohrid Lake Ohrid ( mk, Охридско Езеро , al, Liqeni i Ohrit , also referred as ''Liqeni i Pogradecit'';) is a lake which straddles the mountainous border between the southwestern part of North Macedonia and eastern Albania. It is one of Eu ...

and

Lake Scutari Lake Skadar ( sh-Cyrl-Latn, Скадарско језеро, Skadarsko jezero, ; sq, Liqeni i Shkodrës, ) also called Lake Scutari, Lake Shkodër and Lake Shkodra lies on the border of Albania and Montenegro, and is the largest lake in Southern ...

, and the adjacent Drim River and

Morača River The Morača ( sr-cyrl, Морача, ) is a major river in Montenegro that originates in the northern region in Kolašin Municipality under Mount Rzača. It meanders southwards for before emptying into Lake Skadar. Its drainage basin covers . ...

basins Basin may refer to: Geography and geology * Depression (geology) ** Back-arc basin, a submarine feature associated with island arcs and subduction zones ** Debris basin, designed to prevent damage from debris flow ** Drainage basin (hydrology), a ...

. It is an abundant species in its home range and not considered threatened by the

IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

Footnotes

References

* * (2008)
''Cobitis'' species
Version of 2008-MAY-03. Retrieved 2008-AUG-15. * (2008): The molecular diversity of adriatic spined loaches (Teleostei, Cobitidae). '' Mol. Phylogenet. Evol.'' 46(1): 382–390. (HTML abstract) Cobitis Fish described in 1928 Taxa named by Stanko Karaman Taxonomy articles created by Polbot {{Cobitidae-stub