''Apamea crenata'', known as the clouded-bordered brindle, is a
moth
Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of ...
in the
family
Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Noctuidae
The Noctuidae, commonly known as owlet moths, cutworms or armyworms, are a family of moths. They are considered the most controversial family in the superfamily Noctuoidea because many of the clades are constantly changing, along with the othe ...
. It is distributed throughout the
Palearctic realm
The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa.
The realm consists of several bioregions: the Euro-Si ...
. In the North it crosses the Arctic Circle, in the Mediterranean it is found only in cool locations and mountains avoiding very hot areas. In the Alps, it rises to an altitude of about 2000 metres.
Description
This species has a
wingspan
The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan ...
of 36 to 44 mm. The forewings are pale brown with darker patches along the margins, usually with prominent markings at the base and
tornus. The hindwings are grayish with darker venation.
Melanic forms occur fairly frequently.
Technical description and variation
Forewing pale lilac grey, often washed with pale brownish; the veins paler: costal area tinged with purplish grey; inner margin with white, and with a strong black streak near base: inner and outer lines double, brownish, the outer followed by a double row of dark brown vein dots with a white dot between; claviform stigma brown, with darker outline; orbicular generally elongate and narrow, sometimes shortened and rounder, with centre brown as in reniform and pale outline, the outer half of the reniform paler than the inner, and the lower lobe dark; cell and median shade brown, the latter not reaching below cell; marginal area beyond submarginal line dark redbrown, emitting large teeth inwards on the two folds; hind wing fuscous, pale grey towards base.
Aberration (ab.) ''argentea'' Tutt is a form from Scotland in which the ground colour of forewings is shining silvery whitish, with traces of faint brownish costal streaks; the terminal area and basal streaks pale brown; all other markings obsolete, except a trace of reniform stigma: hindwing pale grey; -
Forma ''ochrea'' Tutt and ''intermedia'' Tutt seem to represent merely the type form when washed with pale ochreous, or with rufous ochreous respectively, this last being the commonest form in Britain; -forma ''flavorufa'' Tutt is a rare form, from the North of England and Scotland only, with ground colour dull yellow red, with the ordinary markings obsolete, except the stigmata which are distinct and outlined with yellow; some short yellow costal dashes, a yellow patch at base of costa and some yellow scales on the dark veins; -the form ''putris'' Hbn. has the whole of forewing suffused with dull grey brown, the median area between inner and outer lines darker brown from costa to inner margin; the white scales of inner margin plainer; some examples are more uniformly dark and with obscured markings, while others are paler brown and show the markings clearly: - forma ''combusta'' Haw. is a dark fuscous form of this, in which the basal area and both hues are mixed with white scales, the outer edge of the reniform being also whitish; ab. ''alopecurus'' Esp.is red-brown with the veins dark, the costa and inner margin and sometimes the veins dusted with whitish, and the stigmata edged with yellow; the two folds redder than the rest; in some cases this red tint predominates and makes the whole wing red; in others the blackish shades overpower the red, these last being ''nigrorubida'' Tutt; - ab. ''subrurea'' Petersen is a form with the forewing darkened with grey brown, with clear markings, and the reniform not picked out with yellow - ''extincta'' Stgr. from
Issykkul and Kuku Nor, has the forewing pale uniform lilac grey; the terminal area and the cell dull redbrown; the hues very indistinctly expressed; the stigmata pale grey, the outer edge of the reniform whitish; the dark form corresponding to this, (as ''alopecurus'' does to typical ''rurea'') - ab. ''uniformis'' ab. nov.
arrenis wholly dull dark liver colour, with no markings except the pale inner and outer lines and the whitish outer edge of the reniform stigma; the hindwing, as in extincta, smooth pale grey.
Biology
''Apamea crenata'' occupies varied
habitats
In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
. It colonizes grass-rich, uncut, and moist to mesophilic places such as wet meadows, fens, forest edges or clearings, mountain and valley meadows, in heaths and moorland areas as well as in gardens and parks.
In the British Isles the moth flies at night from May to July. It is attracted to light and sugar, particularly
nectar-rich
flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanism ...
s.
Adult caterpillars are brown-grey to earth-grey coloured. They have a yellow-and-white dorsal line. On each body segment there are black spots and black point warts. The lateral stripe is yellow and white. Head, the neck shield and anus shield are black-brown. Three bright longitudinal stripes stand out from the neck shield. The maroon pupa is equipped with two thorns and four short hooks on the cremaster. The
larva
A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle.
Th ...
feeds on various
grass
Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in law ...
es including
orchard grasses,
tussock grasses,
fescues, and
canarygrasses. This species overwinters as a larva and feeds in mild weather throughout the season.
References
Further reading
*Chinery, M. ''Collins Guide to the Insects of Britain and Western Europe''. 1986 (Reprinted 1991)
*Skinner, B. ''Colour Identification Guide to Moths of the British Isles''. 1984
External links
Clouded-bordered Brindle at UK MothsTaxonomy
Lepiforum.deVlindernet.nl
{{Taxonbar, from=Q1377434
Apamea (moth)
Moths described in 1766
Palearctic Lepidoptera
Taxa named by Johann Siegfried Hufnagel