''Clintonia umbellulata'', commonly known as white clintonia or speckled wood-lily, is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

in the lily family

Liliaceae The lily family, Liliaceae, consists of about 15 genera and 610 species of flowering plants within the order Liliales. They are monocotyledonous, perennial, herbaceous, often bulbous geophytes. Plants in this family have evolved with a fair ...

. The

specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...

''umbellulata'' means "umbelled," which refers to the shape of the plant’s

inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...

Description

''Clintonia umbellulata'' is a

perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...

herbaceous plant Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...

that spreads by means of underground

rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...

s. A plant stands tall with 2–4 dark green leaves, each long and wide. The inflorescence is a single terminal

umbel In botany, an umbel is an inflorescence that consists of a number of short flower stalks (called pedicels) that spread from a common point, somewhat like umbrella ribs. The word was coined in botanical usage in the 1590s, from Latin ''umbella'' "p ...

with 10–25(–30) outward-facing flowers on a flowering stalk up to high. Each flower has six

tepal A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...

s and six

stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...

s. The tepals are white or greenish white, often marked with purplish brown or green speckles, each tepal being long and wide. The stamens are 60% longer than the tepals. The fruits are black (occasionally ultramarine blue) berries, each long with 2–4 seeds per berry. Each seed is approximately long.

Similar species

Because of their proximity, ''Clintonia umbellulata'' and ''C. borealis'' are often confused. The following table compares the two species character by character (with diagnostic characters emphasized): Based on morphological characters alone, bare-leaved plants may be difficult to identify. In this case, ''Clintonia umbellulata'' is distinguished from ''C. borealis'' by the presence of hairs longer than 1 millimeter on the underside midvein. ''C. umbellulata'' has numerous look-alikes. For example, the inflorescence of the small white leek (''

Allium tricoccum ''Allium tricoccum'' (commonly known as ramp, ramps, ramson, wild leek, wood leek, or wild garlic) is a North American species of wild onion or garlic widespread across eastern Canada and the eastern United States. Many of the common English na ...

'') is very similar in appearance. To distinguish the two, look at the leaves. The leaves of ''A. tricoccum'' have usually wilted by the time the plant is in full bloom while the leaves of ''C. umbellulata'' remain throughout the summer months.

Taxonomy

In 1803,

André Michaux André Michaux, also styled Andrew Michaud, (8 March 174611 October 1802) was a French botanist and explorer. He is most noted for his study of North American flora. In addition Michaux collected specimens in England, Spain, France, and even Per ...

described the species ''Convallaria umbellulata'' Michx., a name that was to become a synonym for ''Clintonia umbellulata'' (Michx.) Morong. The latter was first described by

Thomas Morong Rev. Thomas Morong (April 15, 1827 – April 26, 1894) was an American botanist and clergyman. Biography Morong was born in Cahawba, Alabama to a Massachusetts-born father, Thomas Morong, and a Maryland-born mother Jane Travers. His father owned ...

in 1894. In 1933,

John Kunkel Small John Kunkel Small (January 31, 1869 – January 20, 1938) was an American botanist. Born on January 31, 1869, in Harrisburg Pennsylvania, Kunkel studied botany at Franklin & Marshall College and Columbia University. He was the first Curator of ...

described the segregate species ''Xeniatrum umbellulatum'', a distinction that did not persist. Numerous other synonyms are in use. Perhaps the best known is ''Clintonia alleghaniensis'' Harned, which unlike ''C. umbellulata'' has ultramarine blue (not black) fruit. It is known to occur at a number of sites in

Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth ar ...

Maryland Maryland ( ) is a state in the Mid-Atlantic region of the United States. It shares borders with Virginia, West Virginia, and the District of Columbia to its south and west; Pennsylvania to its north; and Delaware and the Atlantic Ocean to ...

, and

West Virginia West Virginia is a state in the Appalachian, Mid-Atlantic and Southeastern regions of the United States.The Census Bureau and the Association of American Geographers classify West Virginia as part of the Southern United States while the Bur ...

Distribution

''Clintonia umbellulata'' is

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

to the

Appalachian Mountains The Appalachian Mountains, often called the Appalachians, (french: Appalaches), are a system of mountains in eastern to northeastern North America. The Appalachians first formed roughly 480 million years ago during the Ordovician Period. They ...

in the eastern

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

, from

New York New York most commonly refers to: * New York City, the most populous city in the United States, located in the state of New York * New York (state), a state in the northeastern United States New York may also refer to: Film and television * '' ...

Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...

. Counties where the species is known to occur are listed below: * New York: Allegany, Cattaraugus, Chautauqua, Erie, Livingston, Wyoming *

Ohio Ohio () is a state in the Midwestern region of the United States. Of the fifty U.S. states, it is the 34th-largest by area, and with a population of nearly 11.8 million, is the seventh-most populous and tenth-most densely populated. The sta ...

: Ashland, Ashtabula, Columbiana, Coshocton, Harrison, Holmes, Jefferson, Mahoning, Portage, Summit, Trumbull, Tuscarawas, Wayne *

Pennsylvania Pennsylvania (; ( Pennsylvania Dutch: )), officially the Commonwealth of Pennsylvania, is a state spanning the Mid-Atlantic, Northeastern, Appalachian, and Great Lakes regions of the United States. It borders Delaware to its southeast, ...

: Allegheny, Armstrong, Beaver, Bedford, Blair, Butler, Cambria, Centre, Clarion, Clearfield, Crawford, Elk, Erie, Fayette, Forest, Franklin, Fulton, Greene, Huntingdon, Indiana, Jefferson, Lawrence, Mercer, Somerset, Venango, Warren, Washington, Westmoreland *

: Allegany, Garrett *

: Barbour, Braxton, Fayette, Grant, Greenbrier, Hampshire, Hardy, Kanawha, Lincoln, Logan, Marion, McDowell, Mercer, Mineral, Mingo, Monongalia, Monroe, Morgan, Nicholas, Pocahontas, Preston, Randolph, Ritchie, Summers, Tucker, Upshur, Wayne, Webster, Wetzel, Wyoming *

: Albemarle, Alleghany, Amherst, Augusta, Bath, Bedford, Bland, Botetourt, Buchanan, Carroll, Craig, Dickenson, Floyd, Franklin, Giles, Grayson, Greene, Henry, Highland, Lee, Madison, Montgomery, Nelson, Page, Patrick, Pulaski, Rappahannock, Roanoke, Rockbridge, Rockingham, Russell, Scott, Smyth, Tazewell, Warren, Washington, Wise, Wythe *

Kentucky Kentucky ( , ), officially the Commonwealth of Kentucky, is a state in the Southeastern region of the United States and one of the states of the Upper South. It borders Illinois, Indiana, and Ohio to the north; West Virginia and Virginia to ...

: Bell, Breathitt, Clay, Harlan, Jackson, Laurel, Lee, Letcher, Menifee, Morgan, Perry, Powell, Rockcastle, Rowan, Wolf *

Tennessee Tennessee ( , ), officially the State of Tennessee, is a landlocked state in the Southeastern region of the United States. Tennessee is the 36th-largest by area and the 15th-most populous of the 50 states. It is bordered by Kentucky to th ...

: Blount, Campbell, Carter, Cocke, Cumberland, Greene, Hawkins, Johnson, Monroe, Morgan, Polk, Rhea, Sequatchie, Sevier, Sullivan, Unicoi *

North Carolina North Carolina () is a state in the Southeastern region of the United States. The state is the 28th largest and 9th-most populous of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, Georgia and So ...

: Alleghany, Ashe, Avery, Buncombe, Burke, Caldwell, Clay, Graham, Haywood, Henderson, Jackson, Macon, Madison, McDowell, Mitchell, Polk, Rutherford, Surry, Swain, Transylvania, Watauga, Wilkes, Yancey *

South Carolina )''Animis opibusque parati'' ( for, , Latin, Prepared in mind and resources, links=no) , anthem = " Carolina";" South Carolina On My Mind" , Former = Province of South Carolina , seat = Columbia , LargestCity = Charleston , LargestMetro = ...

: Greenville, Oconee, Pickens * Georgia: Gilmer, Habersham, Lumpkin, Murray, Rabun, Towns, Union, White The range of ''C. umbellulata'' apparently overlaps with that of '' C. borealis'' throughout the Appalachian Mountains. (Counties where both species are known to occur are shown above in bold.) Actually ''C. umbellulata'' and ''C. borealis'' are

allopatric Allopatric speciation () – also referred to as geographic speciation, vicariant speciation, or its earlier name the dumbbell model – is a mode of speciation that occurs when biological populations become geographically isolated from ...

, that is, the ranges of the two species do not significantly overlap but are immediately adjacent to one another. ''C. umbellulata'' is globally secure, rare and imperiled in New York, and endangered in Ohio.

References

External links