''Climacoceras'' (from Greek for "

ladder A ladder is a vertical or inclined set of rungs or steps used for climbing or descending. There are two types: rigid ladders that are self-supporting or that may be leaned against a vertical surface such as a wall, and rollable ladders, such a ...

horn Horn most often refers to: *Horn (acoustic), a conical or bell shaped aperture used to guide sound ** Horn (instrument), collective name for tube-shaped wind musical instruments *Horn (anatomy), a pointed, bony projection on the head of various ...

s") is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

artiodactyl The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poster ...

ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, cam ...

s that lived in

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

and

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...

during the

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

. The members of ''Climacoceras'' were related to

giraffe The giraffe is a large African hoofed mammal belonging to the genus ''Giraffa''. It is the tallest living terrestrial animal and the largest ruminant on Earth. Traditionally, giraffes were thought to be one species, ''Giraffa camelopardalis ...

s, and the genus was formerly placed within the

Giraffidae The Giraffidae are a family of ruminant artiodactyl mammals that share a common ancestor with deer and bovids. This family, once a diverse group spread throughout Eurasia and Africa, presently comprises only two extant genera, the giraffe (one or ...

, but is now placed in the

Climacoceratidae Climacoceratidae is a family of superficially deer-like artiodactyl ungulates which lived in the Miocene epoch in Africa. They are close to the ancestry of giraffes, with some genera, such as '' Prolibytherium'', originally classified as giraffes ...

, a sister group within the superfamily

Giraffoidea Giraffoidea is a superfamily that includes the families Climacoceratidae, Antilocapridae, and Giraffidae. The only extant members in the superfamily are the pronghorn, giraffe, and okapi. The Climacoceratidae are also placed in the superfamily, ...

Fossils A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ...

of the two best known

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

of ''Climacoceras'', ''C. africanus'' and ''C. gentryi'', have both been found in

Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...

. The animals measured about tall and had large

ossicone Ossicones are columnar or conical skin-covered bone structures on the heads of giraffes, male okapi, and some of their extinct relatives. Ossicones are distinguished from the superficially similar structures of horns and antlers by their unique ...

s resembling

antlers Antlers are extensions of an animal's skull found in members of the Cervidae (deer) family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally found only on male ...

. ''C. africanus'' had ossicones resembling tall, thorn-covered plant stems, while the ossicones of ''C. gentryi'' resembled thorny

crescent A crescent shape (, ) is a symbol or emblem used to represent the lunar phase in the first quarter (the "sickle moon"), or by extension a symbol representing the Moon itself. In Hinduism, Lord Shiva is often shown wearing a crescent moon on his ...

s. The genus was once placed within

Palaeomerycidae The Palaeomerycidae are an extinct family of ruminants in the order Artiodactyla. Palaeomerycids lived in North America, Europe, Africa and Asia from 33 to 4.9 million years ago (from the Late Eocene to Pliocene epochs), existing for about 28 mi ...

, then Giraffidae. It is now considered a giraffoid in the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Climacoceratidae, erected by Hamilton. Other Climacoceratidae genera include ''

Prolibytherium ''Prolibytherium'' is an extinct genus of prolibytheriid artiodactyl ungulate native to Middle Miocene North Africa and Pakistan, from around 16.9 to 15.97 million years ago. Fossils of ''Prolibytherium'' were found in the Marada Formation of ...

'' from

Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Mediter ...

and

Libya Libya (; ar, ليبيا, Lībiyā), officially the State of Libya ( ar, دولة ليبيا, Dawlat Lībiyā), is a country in the Maghreb region in North Africa. It is bordered by the Mediterranean Sea to the north, Egypt to Egypt–Libya bo ...

and '' Orangemeryx'' from

Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...

and

South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...

References

* MacInnes, D. G. 1936, "A new genus of fossil deer from the Miocene of Africa", Journal of the Linnean Society, Zoology 39: 521–530. * Hamilton, W. R., 1978, "Cervidae and Palaeomerycidae", 495–508, in Maglio, V. J. & Cooke, H. B. S., (eds.) "Evolution of African mammals", Harvard University Press, Cambridge, Massachusetts & London, England, 1978, xiv-641 Climacoceratidae Prehistoric even-toed ungulate genera Miocene even-toed ungulates Miocene mammals of Africa Miocene mammals of Europe Fossil taxa described in 1936 {{paleo-eventoedungulate-stub