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''Claorhynchus'' (meaning "broken
beak The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food ...
", as it is based on broken bones from the snout region) is a
dubious Doubt is a mental state in which the mind remains suspended between two or more contradictory propositions, unable to be certain of any of them. Doubt on an emotional level is indecision between belief and disbelief. It may involve uncertainty ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of cerapodan
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
with a confusing history behind it. It has been considered to be both a
hadrosaur Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which includ ...
id and a ceratopsid, sometimes the same as ''
Triceratops ''Triceratops'' ( ; ) is a genus of herbivore, herbivorous Chasmosaurinae, chasmosaurine Ceratopsidae, ceratopsid dinosaur that first appeared during the late Maastrichtian stage of the Late Cretaceous Period (geology), period, about 68 m ...
'', with two different assignments as to discovery formation and location, and what bones make up its type remains.


History

The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen,
AMNH The American Museum of Natural History (abbreviated as AMNH) is a natural history museum on the Upper West Side of Manhattan in New York City. In Theodore Roosevelt Park, across the street from Central Park, the museum complex comprises 26 i ...
3978, was described by
paleontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
and naturalist
Edward Drinker Cope Edward Drinker Cope (July 28, 1840 – April 12, 1897) was an American zoologist, paleontologist, comparative anatomist, herpetologist, and ichthyologist. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested ...
, who interpreted it as the
rostral bone Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassic. ...
and
predentary Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek st ...
of an "agathaumid" ( ceratopsid) dinosaur, which he said came from the
Laramie Formation The Laramie Formation is a geologic formation of Late Cretaceous (Maastrichtian) age, named by Clarence King in 1876 for exposures in northeastern Colorado, in the United States.King, C. 1876. Report of the Geological Exploration of the Fortieth P ...
of
Colorado Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of t ...
.Cope, E.D. (1892). Fourth note on the Dinosauria of the Laramie. ''The American Naturalist'' 26:756-758. It was soon thought to be a hadrosaurid, though.Hatcher, J.B. (1902). The genus and species of the Trachodontidae (Hadrosauridae, Claosauridae) Marsh. ''Annals of the Carnegie Museum'' 14(1):377-386.Hatcher, J.B., Marsh, O.C., and Lull, R.S. (1907). ''The Ceratopsia''. Government Printing Office:Washington, D.C., 300 pp. In 1904, Franz Baron Nopcsa reclassified it as a ceratopsid. In their influential
monograph A monograph is a specialist work of writing (in contrast to reference works) or exhibition on a single subject or an aspect of a subject, often by a single author or artist, and usually on a scholarly subject. In library cataloging, ''monograph ...
,
Richard Swann Lull Richard Swann Lull (November 6, 1867 – April 22, 1957) was an American paleontologist and Sterling Professor at Yale University who is largely remembered now for championing a non-Darwinian view of evolution, whereby mutation(s) could unl ...
and Nelda E. Wright regarded the genus as a dubious type of hadrosaurid, based on
premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has b ...
e and a
predentary Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek st ...
.Lull, R.S., and Wright, N.E. (1942). ''Hadrosaurian Dinosaurs of North America.'' ''Geological Society of America Special Paper 40'':1-242. This opinion stood until the work of Michael K. Brett-Surman, who stated in his dissertation that, having rediscovered and reexamined the material with Douglas A. Lawson, it was most likely part of a ceratopsid's
neck frill A neck frill is the relatively extensive margin seen on the back of the heads of reptiles with either a bony support such as those present on the skulls of dinosaurs of the suborder Marginocephalia or a cartilaginous one as in the frill-necked ...
, probably part of the
squamosal The squamosal is a skull bone found in most reptiles, amphibians, and birds. In fishes, it is also called the pterotic bone. In most tetrapods, the squamosal and quadratojugal bones form the cheek series of the skull. The bone forms an ancestral co ...
of ''
Triceratops ''Triceratops'' ( ; ) is a genus of herbivore, herbivorous Chasmosaurinae, chasmosaurine Ceratopsidae, ceratopsid dinosaur that first appeared during the late Maastrichtian stage of the Late Cretaceous Period (geology), period, about 68 m ...
''.Brett-Surman, M.K. 1989(1988)
A revision of the Hadrosauridae (Reptilia: Ornithischia) and their evolution during the Campanian and Maastrichtian.
Ph.D. dissertation, George Washington University:Washington, D.C.. pp.1-272.
This information reached
Donald F. Glut Donald F. Glut (; born February 19, 1944) is an American writer, motion picture film director, and screenwriter. He is best known for writing The Empire Strikes Back (novel), the novelization of the second ''Star Wars'' film, ''The Empire Strikes ...
's series of dinosaur
encyclopedia An encyclopedia (American English) or encyclopædia (British English) is a reference work or compendium providing summaries of knowledge either general or special to a particular field or discipline. Encyclopedias are divided into articles ...
s in a confusing form; its entry states that a squamosal and
tooth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
from
South Dakota South Dakota (; Sioux language, Sioux: , ) is a U.S. state in the West North Central states, North Central region of the United States. It is also part of the Great Plains. South Dakota is named after the Lakota people, Lakota and Dakota peo ...
were referred to the genus, and these are what Brett-Surman and Lawson identified, keeping the supposed beak remains separate. Additionally, other major reviews have left the genus as an indeterminate hadrosaurid.Weishampel, D.B., and Horner, J.R. (1990). Hadrosauridae. In: Weishampel, D.B., Dodson, P., and Osmólska, H. (eds.). ''The Dinosauria.'' University of California Press:Berkeley, 534-561. Horner, J.R., Weishampel, D.B., and Forster, C.A. (2004). Hadrosauridae. In: Weishampel, D.B., Dodson, P., and Osmólska, H. (eds.). ''The Dinosauria (second edition)''. University of California Press:Berkeley, 438-463. .


References

{{Taxonbar, from1=Q18606707, from2=Q1006149 Ornithischian genera Late Cretaceous dinosaurs of North America Nomina dubia