Cladophora Ordinata
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''Cladophora'' is a genus of reticulated filamentous
Ulvophyceae The Ulvophyceae or ulvophytes are a class of green algae, distinguished mainly on the basis of ultrastructural morphology, life cycle and molecular phylogenetic data.Graham LE, Graham JM, Wilcox LW (2009) ''Algae''. 2nd Edition. Benjamin Cummin ...
(
green algae The green algae (singular: green alga) are a group consisting of the Prasinodermophyta and its unnamed sister which contains the Chlorophyta and Charophyta/Streptophyta. The land plants (Embryophytes) have emerged deep in the Charophyte alga as ...
). The genus ''Cladophora'' contains many species that are very hard to tell apart and classify, mainly because of the great variation in their appearances, which is affected by habitat, age and environmental conditions. Unlike ''
Spirogyra ''Spirogyra'' (common names include water silk, mermaid's tresses, and blanket weed) is a genus of filamentous charophyte green algae of the order Zygnematales, named for the helical or spiral arrangement of the chloroplasts that is characteris ...
'' the filaments of ''Cladophora'' branch and do not undergo conjugation. There are two multicellular stages in its life cycle – a
haploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respectively ...
gametophyte A gametophyte () is one of the two alternation of generations, alternating multicellular organism, multicellular phases in the life cycles of plants and algae. It is a haploid multicellular organism that develops from a haploid spore that has on ...
and a
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respectively ...
sporophyte A sporophyte () is the diploid multicellular stage in the life cycle of a plant or alga which produces asexual spores. This stage alternates with a multicellular haploid gametophyte phase. Life cycle The sporophyte develops from the zygote pr ...
– which look highly similar. The only way to tell the two stages apart is to either count their
chromosome A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins are ...
s, or examine their offspring. The haploid gametophyte produces haploid gametes by
mitosis In cell biology, mitosis () is a part of the cell cycle in which replicated chromosomes are separated into two new nuclei. Cell division by mitosis gives rise to genetically identical cells in which the total number of chromosomes is mainta ...
and the diploid sporophyte produces haploid spores by
meiosis Meiosis (; , since it is a reductional division) is a special type of cell division of germ cells in sexually-reproducing organisms that produces the gametes, such as sperm or egg cells. It involves two rounds of division that ultimately resu ...
. The only visible difference between the gametes and spores of ''Cladophora'' is that the gametes have two
flagella A flagellum (; ) is a hairlike appendage that protrudes from certain plant and animal sperm cells, and from a wide range of microorganisms to provide motility. Many protists with flagella are termed as flagellates. A microorganism may have f ...
and the spores have four. The ''Cladophora'' species can be a major nuisance causing major alteration to
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
conditions linked particularly with increased
phosphorus Phosphorus is a chemical element with the symbol P and atomic number 15. Elemental phosphorus exists in two major forms, white phosphorus and red phosphorus, but because it is highly reactive, phosphorus is never found as a free element on Ear ...
loading.


Cladophora balls

''Cladophora'' balls are formed from the filaments of the alga which are photosynthetic. Large numbers of these balls were thrown ashore in Devon, England. They had an average diameter of 2.5 cm and several million balls were found forming a layer. This is not to be confused with
marimo Marimo (also known as Cladophora ball, moss ball, or lake ball) is a rare growth form of ''Aegagropila linnaei'' (a species of filamentous green algae) in which the algae grow into large green balls with a velvety appearance. The species can be ...
, which, though formerly part of ''Cladophora'', is now called ''Aegagropila linnaei''.


Laotian Mekong weed

In
Laos Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist ...
, ''Cladophora''
spp. In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate se ...
(ໄຄ "river weed" or more precisely ໄຄຫີນ "rock river weed") are commonly eaten as a delicacy and usually known in English under the name "Mekong weed". The algae grow on underwater rocks and thrive in clear spots of water in the
Mekong The Mekong or Mekong River is a trans-boundary river in East Asia and Southeast Asia. It is the world's List of rivers by length, twelfth longest river and List of longest rivers of Asia, the third longest in Asia. Its estimated length is , ...
river basin. They are harvested 1 to 5 months a year and most often eaten in dry sheets (ໄຄແຜ່ນ
kaipen Kaipen ( lo, ໄຄແຜ່ນ, khai phaen, ) is a Laotian snack made of fresh water green algae, garlic, vegetables, and sesame seeds. ''Kaipen'' is produced in northern Laos and is especially popular in the city of Luang Prabang. During ...
-kháy sheets-), similar to Crispy Chinese Seaweed or Japanese
nori Nori is a dried edible seaweed used in Japanese cuisine, made from species of the red algae genus ''Pyropia'', including ''P. yezonesis'' and '' P. tenera''. It has a strong and distinctive flavor, and is often used to wrap rolls of sushi or '' ...
, though much cruder in their format.
Luang Prabang Luang Phabang, ( Lao: ຫລວງພະບາງ/ ຫຼວງພະບາງ) or ''Louangphabang'' (pronounced ), commonly transliterated into Western languages from the pre-1975 Lao spelling ຫຼວງພຣະບາງ (ຣ = silent r) ...
's speciality is dry khai with sesame (
kaipen Kaipen ( lo, ໄຄແຜ່ນ, khai phaen, ) is a Laotian snack made of fresh water green algae, garlic, vegetables, and sesame seeds. ''Kaipen'' is produced in northern Laos and is especially popular in the city of Luang Prabang. During ...
), while
Vang Vieng Vang may refer to: People Vang is a common surname among Hmong Americans, including *Vang Pao (1929–2011), Lieutenant General in the Royal Lao Army and a leader of the Hmong American community in the United States *Ka Vang (born 1975), writer * ...
is famous for its roasted kháy sheets. They can be eaten in strips as an
appetizer An hors d'oeuvre ( ; french: hors-d'œuvre ), appetiser or starter is a small dish served before a meal in European cuisine. Some hors d'oeuvres are served cold, others hot. Hors d'oeuvres may be served at the dinner table as a part of the ...
, with a meal or as a snack with Beer Lao. Luang Prabang kháy sheets
kaipen Kaipen ( lo, ໄຄແຜ່ນ, khai phaen, ) is a Laotian snack made of fresh water green algae, garlic, vegetables, and sesame seeds. ''Kaipen'' is produced in northern Laos and is especially popular in the city of Luang Prabang. During ...
are the most readily available form of Mekong weed and are famous throughout the country and in the neighbouring
Isaan Northeast Thailand or Isan (Isan/ th, อีสาน, ; lo, ອີສານ; also written as Isaan, Isarn, Issarn, Issan, Esan, or Esarn; from Pali ''īsānna'' or Sanskrit ईशान्य ''īśānya'' "northeast") consists of 20 provin ...
, though difficult to find beyond
Vientiane Vientiane ( , ; lo, ວຽງຈັນ, ''Viangchan'', ) is the capital and largest city of Laos. Vientiane is divided administratively into 9 cities with a total area of only approx. 3,920 square kilometres and is located on the banks of ...
. Mekong weed can also be eaten raw, in soups, or cooked in steamed curries ( lo, ຫມົກໄຄ, ).


''Cladophora'' as an asset and a nuisance

The genus ''Cladophora'' is
cosmopolitan Cosmopolitan may refer to: Food and drink * Cosmopolitan (cocktail), also known as a "Cosmo" History * Rootless cosmopolitan, a Soviet derogatory epithet during Joseph Stalin's anti-Semitic campaign of 1949–1953 Hotels and resorts * Cosmopoli ...
and accordingly infestations cannot often be regarded logically as being invasive. Where they occur they may at various times be seen as beneficial, as a nuisance, or an outright pest. Modest growth of ''Cladophora'' is generally harmless; the growth is an important food for many fish and other aquatic animals, as a buffer for the sequestration of nutrients in the water body and for protection of some aquatic organisms from solar
ultraviolet Ultraviolet (UV) is a form of electromagnetic radiation with wavelength from 10 nanometer, nm (with a corresponding frequency around 30 Hertz, PHz) to 400 nm (750 Hertz, THz), shorter than that of visible light, but longer than ...
radiation. Where ''Cladophora'' becomes a pest is generally where special circumstances cause such drastic overgrowth that
algal bloom An algal bloom or algae bloom is a rapid increase or accumulation in the population of algae in freshwater or marine water systems. It is often recognized by the discoloration in the water from the algae's pigments. The term ''algae'' encompas ...
s develop and form floating mats. Typical examples include where hypertrophication or high mortality of rival organisms produce high concentrations of dissolved phosphorus. Extensive floating mats prevent circulation that is necessary for the aeration of deeper water and, by blocking the light, they kill photosynthesising organisms growing beneath. The mats interfere with the fishing industry by clogging nets and preventing the use of lines. Where they wash ashore the masses of rotting material reduce shoreline property values along water bodies such as the
Great Lakes The Great Lakes, also called the Great Lakes of North America, are a series of large interconnected freshwater lakes in the mid-east region of North America that connect to the Atlantic Ocean via the Saint Lawrence River. There are five lakes ...
in the United States.Great Lakes Science Center,
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Quagga mussel The quagga mussel (''Dreissena rostriformis'', also known as ''Dreissena bugensis'' or ''Dreissena rostriformis bugensis'') is a species (or subspecies) of freshwater mussel, an aquatic bivalve mollusk in the family Dreissenidae. It has an aver ...
populations have increased tremendously during the same time frame as the blooming of ''Cladophora'', though their ecological relationships are not yet clear and may be complex.


Diversity

Species include: *'' Cladophora albida'' *'' Cladophora aokii'' *'' Cladophora brasiliana'' *'' Cladophora catenata'' *'' Cladophora coelothrix'' *''
Cladophora columbiana ''Cladophora'' is a genus of reticulated filamentous Ulvophyceae (green algae). The genus ''Cladophora'' contains many species that are very hard to tell apart and classify, mainly because of the great variation in their appearances, which is a ...
'' *''
Cladophora crispata ''Cladophora'' is a genus of reticulated filamentous Ulvophyceae (green algae). The genus ''Cladophora'' contains many species that are very hard to tell apart and classify, mainly because of the great variation in their appearances, which is ...
'' *'' Cladophora dalmatica'' *'' Cladophora fracta'' *''
Cladophora glomerata ''Cladophora'' is a genus of reticulated filamentous Ulvophyceae (green algae). The genus ''Cladophora'' contains many species that are very hard to tell apart and classify, mainly because of the great variation in their appearances, which is ...
'' *''
Cladophora graminea ''Cladophora'' is a genus of reticulated filamentous Ulvophyceae (green algae). The genus ''Cladophora'' contains many species that are very hard to tell apart and classify, mainly because of the great variation in their appearances, which is ...
'' *'' Cladophora montagneana'' *'' Cladophora ordinata'' *''
Cladophora prolifera ''Cladophora'' is a genus of reticulated Hypha, filamentous Ulvophyceae (green algae). The genus ''Cladophora'' contains many species that are very hard to tell apart and classify, mainly because of the great variation in their appearances, whi ...
'' *'' Cladophora rivularis'' *'' Cladophora rupestris'' *''
Cladophora scopaeformis ''Cladophora'' is a genus of reticulated filamentous Ulvophyceae (green algae). The genus ''Cladophora'' contains many species that are very hard to tell apart and classify, mainly because of the great variation in their appearances, which is ...
'' *'' Cladophora sericea'' *''
Cladophora socialis ''Cladophora socialis'' is a species of green algae. Distribution ''Cladophora socialis'' is found in many locations over the world including the Atlantic Ocean and Pacific Ocean The Pacific Ocean is the largest and deepest of Earth' ...
'' *'' Cladophora vagabunda'' ''
Aegagropila linnaei Marimo (also known as Cladophora ball, moss ball, or lake ball) is a rare growth form of ''Aegagropila linnaei'' (a species of filamentous green algae) in which the algae grow into large green balls with a velvety appearance. The species can be ...
'' (marimo) was formerly placed here as ''Cladophora aegagropila''


References

* *
''Cladophora'' Index.
Monterey Bay Aquarium *Marsin, P. and J. Tomasz. (2005). Introductory studies on the morphology of the genus ''Cladophora'' from the Gulf of Gdańsk. ''Ocean. Hydrob. Studies'', 34(Supl.3): 187-193


External links


Images
of ''Cladophora'' at
AlgaeBase AlgaeBase is a global species database of information on all groups of algae, both marine and freshwater, as well as sea-grass. History AlgaeBase began in March 1996, founded by Michael Guiry. Text was copied from this source, which is avai ...
{{Taxonbar, from=Q311264 Cladophoraceae Cladophorales genera Taxa named by Friedrich Traugott Kützing