Cimolesta is an extinct

order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...

of non-placental

eutheria Eutheria (; from Greek , 'good, right' and , 'beast'; ) is the clade consisting of all therian mammals that are more closely related to placentals than to marsupials. Eutherians are distinguished from noneutherians by various phenotypic tra ...

mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...

s. Cimolestans had a wide variety of body shapes, dentition and lifestyles, though the majority of them were small to medium-sized general mammals that bore superficial resemblances to

rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are na ...

lagomorphs The lagomorphs are the members of the taxonomic order Lagomorpha, of which there are two living families: the Leporidae ( hares and rabbits) and the Ochotonidae (pikas). The name of the order is derived from the Ancient Greek ''lagos'' (λα� ...

mustelid The Mustelidae (; from Latin ''mustela'', weasel) are a family of carnivorous mammals, including weasels, badgers, otters, ferrets, martens, minks and wolverines, among others. Mustelids () are a diverse group and form the largest family in ...

s, and

marsupial Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in a po ...

s. Several groups have previously been suggested to have descended from the Cimolesta: the

Pholidota Pangolins, sometimes known as scaly anteaters, are mammals of the order Pholidota (, from Ancient Greek ϕολιδωτός – "clad in scales"). The one extant family, the Manidae, has three genera: '' Manis'', ''Phataginus'', and ''Smutsia ...

(which would have been regarded as a suborder of Cimolesta), the

Creodonta Creodonta ("meat teeth") is a former order of extinct carnivorous placental mammals that lived from the early Paleocene to the late Miocene epochs in North America, Europe, Asia and Africa. Originally thought to be a single group of animals ance ...

, and the Carnivora. The origins of the enigmatic

Dinocerata Dinocerata (from the Greek (), "terrible", and (), "horn") is an extinct order of plant-eating hoofed mammals with horns and protuberant canine teeth. Classification A 2015 phylogenetic study recovered Dinocerata as closely related to '' ...

have been suggested to lie within the Cimolesta as well. However, recent studies have revealed that cimolestans are more likely to be basal, non-placental eutherians, with no living descendants. Some experts had placed the

pangolin Pangolins, sometimes known as scaly anteaters, are mammals of the order Pholidota (, from Ancient Greek ϕολιδωτός – "clad in scales"). The one extant family, the Manidae, has three genera: '' Manis'', '' Phataginus'', and '' Smut ...

s within Cimolesta, though the current consensus is that the pangolins should be placed within their own order, Pholidota, as a sister taxon to

Carnivora Carnivora is a Clade, monophyletic order of Placentalia, placental mammals consisting of the most recent common ancestor of all felidae, cat-like and canidae, dog-like animals, and all descendants of that ancestor. Members of this group are f ...

within

Ferae Ferae ( , , "wild beasts") is a mirorder of placental mammalsMalcolm C. McKenna, Susan K. Bell: ''Classification of Mammals: Above the Species Level'' in Columbia University Press, New York (1997), 631 Seiten. that groups together clades Pan-Ca ...

. Some have also placed the enigmatic

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Ptolemaiidae Ptolemaiida is a taxon of wolf-sized Afrotheria, afrothere mammals that lived in northern and eastern Africa during the Paleogene. The oldest fossils are from the latest Eocene strata of the Jebel Qatrani Formation, near the Fayum oasis in Egypt. ...

within Cimolesta, also due to similarities between dental and skull anatomies with those of Pantolesta. If the ptolemaiids were indeed cimolestids, then Cimolesta would have ranged from the Late

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...

to the early

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

, when the last ptolemaiid, ''

Kelba Kelba (german: Kelp) is a village in Rapla Parish, Rapla County in northwestern Estonia Estonia, formally the Republic of Estonia, is a country by the Baltic Sea in Northern Europe. It is bordered to the north by the Gulf of Finland ac ...

'', disappeared in Eastern

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

. However, more thorough studies suggest that the ptolemaiids were more probably afrotheres related to

aardvark The aardvark ( ; ''Orycteropus afer'') is a medium-sized, burrowing, nocturnal mammal native to Africa. It is the only living species of the order Tubulidentata, although other prehistoric species and genera of Tubulidentata are known. Unlike ...

tenrec A tenrec is any species of mammal within the afrotherian family Tenrecidae endemic to Madagascar. Tenrecs are wildly diverse; as a result of convergent evolution some resemble hedgehogs, shrews, opossums, rats, and mice. They occupy aquatic, a ...

s and

golden mole Golden moles are small insectivorous burrowing mammals endemic to Sub-Saharan Africa. They comprise the family Chrysochloridae and as such they are taxonomically distinct from the true moles, family Talpidae, and other mole-like families, all o ...

s. If one ignores the ptolemaiids as afrotherians, then the last, unequivocal cimolestids, the pantolestids '' Gobiopithecus'' and '' Kiinkerishella'', died out during the

Late Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "daw ...

or early

Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...

. The cimolestid '' Procerberus'' may have been closely related to

Taeniodonta Taeniodonta ("banded teeth") is an extinct early group of cimolestid mammals known from the Maastrichtian to the Eocene. Taeniodonts evolved quickly into highly specialized digging animals, and varied greatly in size, from rat-sized to species a ...

.Eberle, J. J. (1999). Bridging the transition between didelphodonts and taeniodonts. ''Journal of Paleontology'', ''73'', 936-944.MIDDLETON, M. D. 1983. Early Paleocene vertebrates of the Denver Basin, Colorado. Unpublished Ph.D. dissertation, University of Colorado, Boulder, 404. ''Procerberus'' was the largest cimolestid and different species may be closer to conoryctid taeniodonts and others to stylinodontine taeniodonts. However, ''Procerberus'' has been found by cladistic analysis to be outside of a Taeniodonta + ''Alveugena carbonensis'', with ''Procerberus grandis'' being closer to that clade than to other ''Procerberus'' species.

References

Sources

* * * * {{Taxonbar, from=Q867388 Eocene extinctions Eocene genus extinctions Fossil taxa described in 1975