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Chuckwallas are
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
s found primarily in arid regions of the southwestern United States and northern Mexico. Some are found on coastal islands. The six species of chuckwallas are all placed within the genus ''Sauromalus''; they are part of the iguanid family, Iguanidae.


Taxonomy and etymology

The
generic Generic or generics may refer to: In business * Generic term, a common name used for a range or class of similar things not protected by trademark * Generic brand, a brand for a product that does not have an associated brand or trademark, other ...
name, ''Sauromalus'', is said to be a combination of two ancient Greek words: ''sauros'' meaning "lizard" and ''homalos'' (ὁμαλός) meaning "flat". The common name "chuckwalla" derives from the
Shoshone The Shoshone or Shoshoni ( or ) are a Native American tribe with four large cultural/linguistic divisions: * Eastern Shoshone: Wyoming * Northern Shoshone: southern Idaho * Western Shoshone: Nevada, northern Utah * Goshute: western Utah, easter ...
word ''tcaxxwal'' or Cahuilla ''čaxwal'', transcribed by
Spaniards Spaniards, or Spanish people, are a Romance peoples, Romance ethnic group native to Spain. Within Spain, there are a number of National and regional identity in Spain, national and regional ethnic identities that reflect the country's complex Hist ...
as ''chacahuala''.


Extant species


Description

Chuckwallas are stocky, wide-bodied lizards with flattened midsections and prominent bellies. Their tails are thick, tapering to a blunt tip. Loose folds of skin characterize the neck and sides of their bodies, which are covered in small, coarsely granular scales. The
common chuckwalla ''Sauromalus ater'', also known as the common chuckwalla, is a species of lizard in the family Iguanidae. It inhabits the Sonoran and Mojave Deserts of the Southwestern United States and northwestern Mexico. Its range extends from eastern Cali ...
(''Sauromalus ater'') measures 15 3/4 inches long, whereas insular species such as the San Esteban chuckwalla of San Esteban Island (''Sauromalus varius'') can measure as long as 30 in. They are sexually dimorphic, with males having reddish-pink to orange, yellow, or light gray bodies and black heads, shoulders, and limbs; females and juveniles have bodies with scattered spots or contrasting bands of light and dark in shades of gray or yellow. Males are generally larger than females and possess well-developed femoral pores located on the inner sides of their thighs; these pores produce secretions believed to play a role in marking territory.


Range, habitat, and diet

The genus ''Sauromalus'' has a wide distribution in biomes of the Sonoran and Mojave Deserts. The common chuckwalla ('' S. ater'') is the species with the greatest range, found from southern California east to southern Nevada and Utah and western Arizona, and south to Baja California and northwestern Mexico. The
peninsular chuckwalla ''Sauromalus ater'', also known as the common chuckwalla, is a species of lizard in the family Iguanidae. It inhabits the Sonoran and Mojave Deserts of the Southwestern United States and northwestern Mexico. Its range extends from eastern Cali ...
(''S. australis'') is found on the eastern portion of the southern half of the Baja California Peninsula. The other species are island-dwelling, so have much more restricted distributions. The Angel Island chuckwalla (''S. hispidus'') is found on
Isla Ángel de la Guarda Isla Ángel de la Guarda, (Guardian Angel Island) also called Archangel Island, is a large island in the Gulf of California (Sea of Cortez) east of Bahía de los Ángeles in northwestern Mexico, separated from the Baja California Peninsula by the ...
and surrounding islands off the coast of the Baja California Peninsula. Two rare and
endangered species An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inv ...
are the Montserrat chuckwalla (''S. slevini'') found on Islas Carmen, Coronados, and Montserrat in the southern Gulf of California and the San Esteban chuckwalla or painted chuckwalla (''S. varius'') found on San Esteban Island, Lobos, and Pelicanos. Chuckwallas prefer
lava flow Lava is molten or partially molten rock (magma) that has been expelled from the interior of a terrestrial planet (such as Earth) or a moon onto its surface. Lava may be erupted at a volcano or through a fracture in the crust, on land or und ...
s and rocky areas typically vegetated by creosote bush and other such drought-tolerant scrub. The lizards may be found at elevations up to 4,500 ft (1,370 m). Primarily herbivorous, chuckwallas feed on leaves, fruit, and flowers of annuals and perennial plants; insects represent a supplementary prey. The lizards are said to prefer yellow flowers, such as those of the brittlebush (''Encelia farinosa'').


Behavior and reproduction

Harmless to humans, these lizards are known to run from potential threats. When disturbed, a chuckwalla wedges itself into a tight rock crevice and inflates its lungs to entrench itself.Stebbins, Robert C., (2003) ''A Field Guide to Western Reptiles and Amphibians'', 3rd Edition. Houghton Mifflin Company, Deban, S.M., J.C. O’Reilly, and T.C. Theimer 1994. Mechanism of defensive inflation in the chuckwalla, Sauromalus obesus. Journal of Experimental Zoology 270: 451-459 Males are seasonally and conditionally territorial; an abundance of resources tends to create a hierarchy based on size, with one large male dominating the area's smaller males. Chuckwallas use a combination of color and physical displays, namely "push-ups", head-hobbing, and gaping of the mouth, to
communicate Communication (from la, communicare, meaning "to share" or "to be in relation with") is usually defined as the transmission of information. The term may also refer to the message communicated through such transmissions or the field of inquir ...
and defend their territory. Chuckwallas are diurnal animals and as they are ectothermic, spend much of their mornings and winter days basking. These lizards are well adapted to desert conditions; they are active at temperatures up to . Chuckwallas
hibernate Hibernation is a state of minimal activity and metabolic depression undergone by some animal species. Hibernation is a seasonal heterothermy characterized by low body-temperature, slow breathing and heart-rate, and low metabolic rate. It most ...
during cooler months and emerge in February. Juveniles emerge first, then adults, as temperatures reach around . Mating occurs from April to July, with five to 16 eggs laid between June and August. The eggs hatch in late September. Chuckwallas may live for 25 years or more. File:Male Chuckwalla.JPG, Large male chuckwalla, picture taken in the White Tank Mountains near Surprise, AZ File:Chuckwalla.jpg, Adult chuckwalla of the Sonoran Desert File:Juvenile Chuckwalla.jpg, Juvenile chuckwalla of the Sonoran Desert File:Chuckwalla(landers ca).jpg, Adult chuckwalla of the Mojave Desert File:Basking Chuckwalla.jpg, Large male common chuckwalla in Joshua Tree National Park File:Chuckwalla Sneezing.jpg, A chuckwalla sneezing salt in Anza-Borrego Desert State Park


Human use

The Seri people considered the Angel Island species of chuckwalla an important food item.Richard Felger and Mary B. Moser (1985) ''People of the desert and sea: ethnobotany of the Seri Indians'' Tucson: University of Arizona Press. They are believed to have translocated the lizards to most of the islands in Bahia de los Angeles for use as a food source in times of need.


References


External links

*ARKive
images and movies of the San Esteban Island chuckwalla ''(Sauromalus varius)''
www.chuckwalla-reptiles-tirol.at
Sauromalus ( Chuckwalla ) and Crotaphytus
{{Taxonbar, from=Q2016288 Reptiles of Mexico Fauna of the Colorado Desert Reptiles of the United States Taxa named by Auguste Duméril sv:Chuckwalla