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Chromodorididae, or chromodorids, are a
taxonomic Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
of colourful, sea slugs; dorid nudibranchs,
marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military * ...
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is e ...
s in the superfamily Doridoidea. “Chromodorid nudibranchs are among the most gorgeously coloured of all animals.” The over 360 described species are primarily found in tropical and subtropical waters, as members of coral reef communities, specifically associated with their sponge prey. The chromodorids are the most speciose family of opisthobranchs. They range in size from <10mm to over 30 cm, although most species are approximately 15–30 mm in size. Although, they have a worldwide distribution, most species are found in the
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
region. A scientific paper published in 2007, found the most widespread chromodorid genera, (''
Mexichromis ''Mexichromis'' is a genus of colourful sea slugs, dorid nudibranchs, shell-less marine gastropod mollusks in the family Chromodorididae.Caballer, M.; Bouchet, P. (2011)''Mexichromis'' Bertsch, 1977.Accessed through: World Register of Marine Spe ...
'', ''
Chromodoris ''Chromodoris'' is a genus of very colourful sea slugs or dorid nudibranchs, marine gastropod molluscs, and the type genus of the family Chromodorididae. Within the genus ''Chromodoris'', there are currently 101 classified species. Species within ...
'', ''
Glossodoris ''Glossodoris'' is a genus of sea slugs, dorid nudibranchs, shell-less marine gastropod mollusks in the family Chromodorididae.Caballer, M.; Gofas, S. (2012)''Glossodoris'' Ehrenberg, 1831.Accessed through: World Register of Marine Species on 201 ...
'' and ''
Hypselodoris ''Hypselodoris'' is a genus of colourful sea slugs or dorid nudibranchs, marine gastropod mollusks in the family Chromodorididae.MolluscaBase (2018)''Hypselodoris'' Stimpson, 1855.Accessed on 2018-09-16. Modern usage follows a more restricted v ...
'') to be
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
or polyphyletic. The family Cadlinidae
Bergh Bergh is a former municipality in the Dutch province of Gelderland. The municipality included the villages of 's-Heerenberg, Zeddam, and Stokkum. In 2005, it merged with Didam to form the new municipality of Montferland Montferland () is ...
, 1891
has been considered a synonym of the Chromodorididae." Research by R.F. Johnson in 2011 has shown that ''
Cadlina ''Cadlina'' is a genus of sea slugs, dorid nudibranchs, shell-less marine gastropod mollusks historically classified in the family Chromodorididae.Bouchet, P.; Gofas, S. (2011). Cadlina Bergh, 1878. Accessed through: World Register of Marine Spec ...
'' does not belong to the family Chromodorididae. She has therefore brought back the name
Cadlinidae Cadlinidae is a family of sea slugs, dorid nudibranchs, marine gastropod mollusks in the superfamily Doridoidea. Molecular phylogenetic studies and their taxon-sampling schemes can have a strong influence of the resulting phylogeny. Research by ...
from synonymy with ''Chromodorididae''. The chromodorid nudibranchs without ''Cadlina'' are now
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
and turn out to be a possible sister to the family
Actinocyclidae Actinocyclidae is a family of sea slugs, dorid nudibranchs, marine gastropod mollusks in the superfamily Doridoidea. This family is within the clade Euctenidiacea. Taxonomy Genera within the family Actinocyclidae include: * ''Actinocyclus'' E ...
. ''Cadlina'' and ''
Aldisa Aldisa is a genus of sea slugs, dorid nudibranchs, marine gastropod molluscs in the family Cadlinidae.Valdés Á. (2002). ''A phylogenetic analysis and systematic revision of the cryptobranch dorids (Mollusca, Nudibranchia, Anthobranchia)''. Zool ...
'' are the only two genera currently classified in the Cadlinidae. A comprehensive phylogeny of the chromodorid nudibranchs found every one of the 14 traditional chromodorid genera were either non-monophyletic, or rendered another genus paraphyletic. Additionally, both the monotypic genera ''Verconia'' and ''Diversidoris'' are nested within clades. The authors presented a new classification of the chromodorid nudibranchs, which used molecular data to untangle evolutionary relationships and at the same time retains a historical connection to traditional systematics by using generic names attached to
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
as clade names. All Chromodorid nudibranchs feed on
sponges Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate through ...
.Rudman W.B. & Bergquist, P.R., 2007. ''A review of feeding specificity in the sponge-feeding Chromodorididae (Nudibranchia: Mollusca).'' Molluscan Research 27(2): 60–88.


Genera

Genera within the family Chromodorididae include: * ''
Ardeadoris ''Ardeadoris'' is a genus of sea slugs, specifically dorid nudibranchs, shell-less marine gastropod mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs ...
'' Rudman, 1984 * ''
Berlanguella ''Berlanguella'' is a genus of sea slugs, specifically dorid nudibranchs, shell-less marine gastropod molluscs in the family Chromodorididae.Bouchet, P. (2011)''Berlanguella'' Ortea, Bacallado & Valdés, 1992, accessed through World Register of M ...
'' Ortea, Bacallado & Valdés, 1992 (although this has not been properly investigated) * ''
Cadlinella ''Cadlinella'' is a genus of colourful sea slugs, specifically dorid nudibranchs, shell-less marine gastropod mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as ...
'' Thiele, 1931 * '' Ceratosoma'' J. E. Gray and M. E. Gray, 1850 * ''
Chromodoris ''Chromodoris'' is a genus of very colourful sea slugs or dorid nudibranchs, marine gastropod molluscs, and the type genus of the family Chromodorididae. Within the genus ''Chromodoris'', there are currently 101 classified species. Species within ...
'' Alder and Hacock, 1855 - the type genus * ''
Diversidoris ''Diversidoris'' is a genus of sea slugs, dorid nudibranchs, shell-less marine gastropod mollusks in the family Chromodorididae.MolluscaBase (2018)''Diversidoris'' Rudman, 1987.Accessed on 2019-01-31. This clade is supported by molecular phylogen ...
'' Rudman, 1987 * ''
Doriprismatica ''Doriprismatica'' is a genus of sea slugs, dorid nudibranchs, shell-less marine gastropod mollusks in the family Chromodorididae Chromodorididae, or chromodorids, are a taxonomic family of colourful, sea slugs; dorid nudibranchs, marine gas ...
'' d'Orbigny, 1839 * ''
Durvilledoris ''Durvilledoris'' was a genus of sea slugs, dorid nudibranchs, shell-less marine gastropod mollusks in the family Chromodorididae. This genus has become a synonym of ''Mexichromis'' Bertsch, 1977 Bouchet, P.; Caballer, M. (2012)''Durvilledoris'' ...
'' Rudman, 1984 * ''
Felimare ''Felimare'' is a genus of sea slugs, dorid nudibranchs, shell-less marine gastropod molluscs in the subfamily Miamirinae of the family Chromodorididae.Bouchet, P.; Caballer, M. (2012)''Felimare''.Accessed through: World Register of Marine Speci ...
'' Ev. Marcus & Er. Marcus, 1967 * ''
Felimida ''Felimida'' is a genus of sea slugs, dorid nudibranchs, shell-less marine gastropod mollusks in the family Chromodorididae.Bouchet, P.; Caballer, M. (2012). ''Felimida''. Accessed through: World Register of Marine Species at http://www.marinespe ...
'' Marcus, 1971 * ''
Glossodoris ''Glossodoris'' is a genus of sea slugs, dorid nudibranchs, shell-less marine gastropod mollusks in the family Chromodorididae.Caballer, M.; Gofas, S. (2012)''Glossodoris'' Ehrenberg, 1831.Accessed through: World Register of Marine Species on 201 ...
'' Ehrenberg, 1831 * ''
Goniobranchus ''Goniobranchus'' is a genus of sea slugs, dorid nudibranchs, shell-less marine gastropod molluscs in the family Chromodorididae.Bouchet, P.; Caballer, M. (2012)''Goniobranchus''.Accessed through: World Register of Marine Species on 2012-04-22 ...
'' Pease, 1866 * ''
Hypselodoris ''Hypselodoris'' is a genus of colourful sea slugs or dorid nudibranchs, marine gastropod mollusks in the family Chromodorididae.MolluscaBase (2018)''Hypselodoris'' Stimpson, 1855.Accessed on 2018-09-16. Modern usage follows a more restricted v ...
'' Stimpson, 1855 * ''
Mexichromis ''Mexichromis'' is a genus of colourful sea slugs, dorid nudibranchs, shell-less marine gastropod mollusks in the family Chromodorididae.Caballer, M.; Bouchet, P. (2011)''Mexichromis'' Bertsch, 1977.Accessed through: World Register of Marine Spe ...
'' Bertsch, 1977 * ''
Miamira ''Miamira'' is a genus of colourful sea slugs, specifically dorid nudibranchs, shell-less marine gastropod mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as mo ...
'' Bergh, 1874 * ''
Thorunna ''Thorunna'' is a genus of sea slugs, dorid nudibranchs, shell-less marine (ocean), marine gastropod mollusks in the family Chromodorididae. Description The genus ''Thorunna'' contains chromodorid nudibranchs with a distinctive radula, which is ...
''
Bergh Bergh is a former municipality in the Dutch province of Gelderland. The municipality included the villages of 's-Heerenberg, Zeddam, and Stokkum. In 2005, it merged with Didam to form the new municipality of Montferland Montferland () is ...
, 1878
* '' Tyrinna''
Bergh Bergh is a former municipality in the Dutch province of Gelderland. The municipality included the villages of 's-Heerenberg, Zeddam, and Stokkum. In 2005, it merged with Didam to form the new municipality of Montferland Montferland () is ...
, 1898
* ''
Verconia ''Verconia'' is a genus of sea slugs, dorid nudibranchs, shell-less marine (ocean), marine gastropod mollusks in the family Chromodorididae.MolluscaBase (2018)''Verconia'' Pruvot-Fol, 1931.Accessed on 2018-12-25. The host sponges for species in ...
'' Pruvot-Fol, 1931 - synonym: '' Noumea'' Risbec, 1928 ; Genera brought into synonymy : * ''Actinodoris'' Ehrenberg, 1831 : synonym of ''
Chromodoris ''Chromodoris'' is a genus of very colourful sea slugs or dorid nudibranchs, marine gastropod molluscs, and the type genus of the family Chromodorididae. Within the genus ''Chromodoris'', there are currently 101 classified species. Species within ...
'' Alder & Hancock, 1855 * ''Babaina'' Odhner in Franc, 1968 : synonym of ''
Thorunna ''Thorunna'' is a genus of sea slugs, dorid nudibranchs, shell-less marine (ocean), marine gastropod mollusks in the family Chromodorididae. Description The genus ''Thorunna'' contains chromodorid nudibranchs with a distinctive radula, which is ...
'' Bergh, 1878 * ''Casella'' H. Adams & A. Adams, 1854 : synonym of ''
Glossodoris ''Glossodoris'' is a genus of sea slugs, dorid nudibranchs, shell-less marine gastropod mollusks in the family Chromodorididae.Caballer, M.; Gofas, S. (2012)''Glossodoris'' Ehrenberg, 1831.Accessed through: World Register of Marine Species on 201 ...
'' Ehrenberg, 1831 * ''Chromolaichma'' Bertsch, 1977 : synonym of ''
Glossodoris ''Glossodoris'' is a genus of sea slugs, dorid nudibranchs, shell-less marine gastropod mollusks in the family Chromodorididae.Caballer, M.; Gofas, S. (2012)''Glossodoris'' Ehrenberg, 1831.Accessed through: World Register of Marine Species on 201 ...
'' Ehrenberg, 1831 * ''Crepidodoris'' Pagenstecher, 1877 : synonym of ''
Glossodoris ''Glossodoris'' is a genus of sea slugs, dorid nudibranchs, shell-less marine gastropod mollusks in the family Chromodorididae.Caballer, M.; Gofas, S. (2012)''Glossodoris'' Ehrenberg, 1831.Accessed through: World Register of Marine Species on 201 ...
'' Ehrenberg, 1831 * ''Digidentis'' Rudman, 1984 : synonym of ''
Thorunna ''Thorunna'' is a genus of sea slugs, dorid nudibranchs, shell-less marine (ocean), marine gastropod mollusks in the family Chromodorididae. Description The genus ''Thorunna'' contains chromodorid nudibranchs with a distinctive radula, which is ...
'' * ''Jeanrisbecia'' Franc, 1968 : synonym of ''
Risbecia ''Risbecia'' is a genus of colorful sea slugs, dorid nudibranchs, shell-less marine gastropod mollusks in the family Chromodorididae.Caballer, M. (2011). Risbecia Odhner, 1934. Accessed through: World Register of Marine Species at http://www.m ...
'' Odhner, 1934 synonym of ''
Hypselodoris ''Hypselodoris'' is a genus of colourful sea slugs or dorid nudibranchs, marine gastropod mollusks in the family Chromodorididae.MolluscaBase (2018)''Hypselodoris'' Stimpson, 1855.Accessed on 2018-09-16. Modern usage follows a more restricted v ...
'' Stimpson, 1855 * ''Lissodoris'' Odhner, 1934 : synonym of ''
Chromodoris ''Chromodoris'' is a genus of very colourful sea slugs or dorid nudibranchs, marine gastropod molluscs, and the type genus of the family Chromodorididae. Within the genus ''Chromodoris'', there are currently 101 classified species. Species within ...
'' Alder & Hancock, 1855 * ''Noumea'' (preoccupied by a beetle): synonym of ''
Verconia ''Verconia'' is a genus of sea slugs, dorid nudibranchs, shell-less marine (ocean), marine gastropod mollusks in the family Chromodorididae.MolluscaBase (2018)''Verconia'' Pruvot-Fol, 1931.Accessed on 2018-12-25. The host sponges for species in ...
'' * ''Orodoris'' Bergh, 1875 : synonym of '' Ceratosoma'' Gray, 1850 * ''Pectenodoris'' Rudman, 1984 : synonym of ''
Mexichromis ''Mexichromis'' is a genus of colourful sea slugs, dorid nudibranchs, shell-less marine gastropod mollusks in the family Chromodorididae.Caballer, M.; Bouchet, P. (2011)''Mexichromis'' Bertsch, 1977.Accessed through: World Register of Marine Spe ...
'' * ''Pterodoris'' Ehrenberg, 1831 : synonym of ''
Hypselodoris ''Hypselodoris'' is a genus of colourful sea slugs or dorid nudibranchs, marine gastropod mollusks in the family Chromodorididae.MolluscaBase (2018)''Hypselodoris'' Stimpson, 1855.Accessed on 2018-09-16. Modern usage follows a more restricted v ...
'' Stimpson, 1855 * ''Risbecia'' Odhner, 1934 : synonym of ''
Hypselodoris ''Hypselodoris'' is a genus of colourful sea slugs or dorid nudibranchs, marine gastropod mollusks in the family Chromodorididae.MolluscaBase (2018)''Hypselodoris'' Stimpson, 1855.Accessed on 2018-09-16. Modern usage follows a more restricted v ...
'' * ''Rosodoris'' Pruvot-Fol, 1954 : synonym of ''
Glossodoris ''Glossodoris'' is a genus of sea slugs, dorid nudibranchs, shell-less marine gastropod mollusks in the family Chromodorididae.Caballer, M.; Gofas, S. (2012)''Glossodoris'' Ehrenberg, 1831.Accessed through: World Register of Marine Species on 201 ...
'' Ehrenberg, 1831 Image:Glossodoris cruenta 3.jpg, ''
Ardeadoris cruenta ''Ardeadoris cruenta'' is a species of very colourful sea slug, a dorid nudibranch, a marine gastropod mollusc in the family Chromodorididae.Bouchet, P. (2012)''Ardeadoris cruenta''.Accessed through: World Register of Marine Species on 2012-06- ...
'' Image:Ceratosoma tenue - Purple Edged Ceratosoma Nudibranch (2649065671).jpg, ''
Ceratosoma tenue is a species of colourful dorid nudibranch, a sea slug, a shell-less marine gastropod mollusc in the family Chromodorididae. Distribution This sea slug is widespread throughout the tropical waters of the Indo-Pacific area, from the oriental ...
'' Image:Chromodoris magnifa.JPG, '' Chromodoris magnifica'' Image:Nudibranch - Noumea crocea 3.jpg, ''
Diversidoris crocea ''Diversidoris crocea'' is a species of colourful sea slug, a dorid nudibranch, a shell-less marine gastropod mollusk in the family Chromodorididae.MolluscaBase (2018)''Diversidoris crocea'' (Rudman, 1986).Accessed on 2019-01-31. Distribution ...
'' Image:Glossodoris atromarginata.jpg, ''
Doriprismatica atromarginata ''Doriprismatica atromarginata'' is a species of sea slug. It is a dorid nudibranch, a shell-less marine gastropod mollusk in the family Chromodorididae.MolluscaBase (2018)''Doriprismatica atromarginata'' (Cuvier, 1804).Accessed on 2018-12-25. ...
'' Image:Nudibrankioa.jpg, ''
Felimare cantabrica ''Felimare cantabrica'' is a species of colourful sea slug or dorid nudibranch, a marine gastropod mollusc in the family Chromodorididae.Bouchet, P. (2012)''Felimare cantabrica''.Accessed through: World Register of Marine Species on 2012-05-11 ...
'' Image:Chromodoris krohni 2 (Vérany, 1846) .jpg, ''
Felimida krohni ''Felimida krohni'' is a species of colorful sea slug, a dorid nudibranch, a marine gastropod mollusc in the family Chromodorididae.Bouchet, P. (2012). Felimida krohni. Accessed through: World Register of Marine Species at http://www.marinespec ...
'' Image:Glossodoris cincta.jpg, ''
Glossodoris cincta ''Glossodoris cincta'' is a species of sea slug, a dorid nudibranch, a shell-less marine gastropod mollusk in the family Chromodorididae. This name has been extensively used for a group of similarly coloured species, some of which are un-named. ...
'' Image:ChromodorisGeminus.jpg, ''
Goniobranchus geminus ''Goniobranchus geminus'', also known as the gem sea slug, is a species of very colourful sea slug, a dorid nudibranch, a marine gastropod mollusc in the family Chromodorididae.Bouchet, P. (2012)''Goniobranchus geminus''.Accessed through: World R ...
'' Image:Hypselodoris infucata.jpg, ''
Hypselodoris confetti ''Hypselodoris confetti'' is a species of sea slug or dorid nudibranch, a marine gastropod mollusc in the family Chromodorididae. Distribution This nudibranch was described from Siar Island, Madang, Papua New Guinea, . It is reported from Indone ...
'' Image:Mexichromis multituberculata 1.jpg, ''
Mexichromis multituberculata ''Mexichromis multituberculata'' is a species of sea slug, a dorid nudibranch, a shell-less marine gastropod mollusk in the family Chromodorididae. Distribution This animal was initially found in the Seto Inland Sea and Kii Channel, Japan ...
'' Image:Noumea norba.jpg, ''
Verconia norba ''Verconia norba'' is a species of colorful sea slug, a dorid nudibranch, a shell-less marine gastropod mollusk in the family Chromodorididae.Bieler, R. (2015). Verconia norba (Er. Marcus & Ev. Marcus, 1970). In: MolluscaBase (2015). Accessed ...
'' Image:Thorunna australis003.jpg, ''
Thorunna australis ''Thorunna australis'' is a species of sea slug, a dorid nudibranch, a shell-less marine gastropod mollusk in the family Chromodorididae.MolluscaBase (2018)''Thorunna australis'' (Risbec, 1928).Accessed on 2018-12-23. Distribution This spec ...
''


References

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