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The chilling requirement of a fruit is the minimum period of cold
weather Weather is the state of the atmosphere, describing for example the degree to which it is hot or cold, wet or dry, calm or stormy, clear or cloud cover, cloudy. On Earth, most weather phenomena occur in the lowest layer of the planet's atmos ...
after which a
fruit-bearing tree A fruit tree is a tree which bears fruit that is consumed or used by animals and humans — all trees that are flowering plants produce fruit, which are the ripened ovaries of flowers containing one or more seeds. In horticultural usage, t ...
will blossom. It is often expressed in chill hours, which can be calculated in different ways, all of which essentially involve adding up the total amount of time in a winter spent at certain temperatures. Some bulbs have chilling requirements to bloom, and some seeds have chilling requirements to sprout. Biologically, the chilling requirement is a way of ensuring that
vernalization Vernalization (from Latin ''vernus'', "of the spring") is the induction of a plant's flowering process by exposure to the prolonged cold of winter, or by an artificial equivalent. After vernalization, plants have acquired the ability to flower, ...
occurs.


Chilling units or chilling hours

A ''chilling unit'' in
agriculture Agriculture or farming is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled people t ...
is a metric of a plant's exposure to chilling temperatures. Chilling temperatures extend from
freezing point The melting point (or, rarely, liquefaction point) of a substance is the temperature at which it changes state from solid to liquid. At the melting point the solid and liquid phase exist in equilibrium. The melting point of a substance depend ...
to, depending on the model, or even .Byrne, D. H., and T. A. Bacon (1992).
Chilling estimation: its importance and estimation
''. The Texas Horticulturist 18(8):5, 8-9. Retrieved 2010-05-24.
Stone fruit In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part ( exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or ''pyrena'') of hardened endocarp with a seed (''kernel' ...
trees and certain other plants of
temperate climate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout ...
develop next year's
bud In botany, a bud is an undeveloped or embryonic shoot and normally occurs in the axil of a leaf or at the tip of a stem. Once formed, a bud may remain for some time in a dormant condition, or it may form a shoot immediately. Buds may be spec ...
s in the summer. In the autumn the buds become dormant, and the switch to proper, healthy dormancy is triggered by a certain minimum exposure to chilling temperatures. Lack of such exposure results in delayed and substandard foliation, flowering and fruiting. One chilling unit, in the simplest models, is equal to one hour's exposure to the chilling temperature; these units are summed up for a whole season. Advanced models assign different weights to different temperature bands.


Requirements

According to Fishman, chilling in trees acts in two stages. The first is reversible: chilling helps to build up the
precursor Precursor or Precursors may refer to: * Precursor (religion), a forerunner, predecessor ** The Precursor, John the Baptist Science and technology * Precursor (bird), a hypothesized genus of fossil birds that was composed of fossilized parts of u ...
to dormancy, but the process can be easily reversed with a rise in temperature. After the level of precursor reaches a certain threshold, dormancy becomes irreversible and will not be affected by short-term warm temperature peaks.
Apple An apple is an edible fruit produced by an apple tree (''Malus domestica''). Apple trees are cultivated worldwide and are the most widely grown species in the genus ''Malus''. The tree originated in Central Asia, where its wild ancestor, ' ...
s have the highest chilling requirements of all fruit trees, followed by apricots and, lastly,
peach The peach (''Prunus persica'') is a deciduous tree first domesticated and cultivated in Zhejiang province of Eastern China. It bears edible juicy fruits with various characteristics, most called peaches and others (the glossy-skinned, non-f ...
es. Apple
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s have a diverse range of permissible minimum chilling: most have been bred for temperate weather, but ''
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'' and ''
Fuji Fuji may refer to: Places China * Fuji, Xiangcheng City (付集镇), town in Xiangcheng City, Henan Japan * Mount Fuji, the tallest mountain in Japan * Fuji River * Fuji, Saga, town in Saga Prefecture * Fuji, Shizuoka, city in Shizuoka Prefec ...
'' can be successfully grown in subtropical Bakersfield, California.Hall, Anthony (2001).
Crop responses to environment
'. CRC Press. . p. 87.
Peach cultivars in
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range in their requirements from 100 chilling units (''Florida Grande'' cultivar, zoned for low chill regions) to 1,000 units (''Surecrop'', zoned for high chill regions).Kamas, J.; McEachern, J. R; Stein, L.; Roe, N. (1998).
Peach Production in Texas
', table 1. Texas A&M University. Retrieved 2010-05-24.
Planting a low-chilling cultivar in a high-chill region risks loss of a year's harvest when an early bloom is hit by a spring frost. A high-chilling cultivar planted in a low-chill region will, quite likely, never fruit at all. A four-year study of ''Ruston Red''
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peach, which has a threshold of 850 chilling units, demonstrated that a seasonal chilling deficiency of less than 50 units has no effect on harvest. Deficiency of 50 to 100 units may result in loss of up to 50% of expected harvest. Deficiency of 250 hours and more is a sure loss of practically whole harvest; the few fruit will be of very poor quality and have no market value.Powell, A. (1999).
Action Program for Dormex Application on Peaches
''. Auburn University. Retrieved 2010-05-24.
Rest-breaking agents (e.g. hydrogen cyanamide, trade name ''BudPro or Dormex''), applied in spring, can partially mitigate the effects of insufficient chilling. BudPro can substitute for up to 300 hours of chilling, but an excessive spraying and timing error can easily damage the buds. Other products such as Dormex use stabilizing compounds. Chilling of orange trees has two effects. First, it increases production of carotenoids and decreases chlorophyll content of the fruit, improving their appearance and, ultimately, their market value. Second, the "quasi-dormancy" experienced by orange trees triggers concentrated flowering in spring, as opposed to more or less uniform round-the-year flowering and fruiting in warmer climates. Biennial plants like cabbage, sugar beet,
celery Celery (''Apium graveolens'') is a marshland plant in the family Apiaceae that has been cultivated as a vegetable since antiquity. Celery has a long fibrous stalk tapering into leaves. Depending on location and cultivar, either its stalks, ...
and carrots need chilling to develop second-year flowering buds. Excessive chilling in the early stages of a sugar beet seedling, on the contrary, may trigger undesired growth of a flowering stem ( bolting) in its first year. This phenomenon has been offset by breeding sugar beet cultivars with a higher minimum chilling threshold. Such cultivars can be seeded earlier than normal without the risk of bolting.


Models

All models require hourly recording of temperatures. The simplest model assigns one chilling unit for every full hour at temperatures below . A slightly more sophisticated model excludes freezing temperatures, which do not contribute to proper dormancy cycle, and counts only hours with temperatures between and . The Utah model assigns different weight to different temperature bands; a full unit per hour is assigned only to temperatures between and . Maximum effect is achieved at . Temperatures between and (the threshold between chilling and warm weather) have zero weight, and higher temperature have negative weights: they reduce the beneficial effects of an already accumulated chilling hours. Southwick et al. wrote that neither of these models is accurate enough to account for application of rest-breaking agents widely used in modern farming. They advocated the use of a dynamic model tailored to the two-stage explanation of dormancy.Southwick, S.; Khan, Z.; Glozer, K. (2003).
Evaluation of Chill Models from Historical Rest-Breaking Spray Experiments on Bing Sweet Cherry
'.
University of California, Davis The University of California, Davis (UC Davis, UCD, or Davis) is a public land-grant research university near Davis, California. Named a Public Ivy, it is the northernmost of the ten campuses of the University of California system. The inst ...
. Retrieved 2010-05-24.


References


External links


Harvest Prediction Model for the counties and towns of California
University of California The University of California (UC) is a public land-grant research university system in the U.S. state of California. The system is composed of the campuses at Berkeley, Davis, Irvine, Los Angeles, Merced, Riverside, San Diego, San Franci ...
Agricultural and Natural Resources.
gardenweb discussion of chilling requirementspermaculture discussion of chilling requirements
{{Webarchive, url=https://web.archive.org/web/20130114152336/http://phoenixpermaculture.ning.com/forum/topics/winter-chill-data-for-phoenix?commentId=2008067%3AComment%3A156520 , date=2013-01-14
chill accumulation calculator for wunderground.com weather stations
Horticulture Meteorological indices Plant physiology