Chilean Hawk
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The Chilean hawk (''Accipiter chilensis'') is a
bird of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predators h ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
belonging to the typical hawks. It breeds in
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
forests from central
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
and western
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
south to
Tierra del Fuego Tierra del Fuego (, ; Spanish for "Land of the Fire", rarely also Fireland in English) is an archipelago off the southernmost tip of the South American mainland, across the Strait of Magellan. The archipelago consists of the main island, Isla G ...
, from sea level to 2,700 m altitude (though birds are rarely observed above 1,000 m). Some
winter Winter is the coldest season of the year in polar and temperate climates. It occurs after autumn and before spring. The tilt of Earth's axis causes seasons; winter occurs when a hemisphere is oriented away from the Sun. Different cultures ...
apparently in the lowlands of NW Argentina. It is sometimes considered to be a subspecies of its northern relative the
bicolored hawk The bicolored hawk (''Accipiter bicolor'') is a species of bird of prey in the family Accipitridae. It is found in forest, woodland, second growth, plantations, and wooded savanna in southeastern Mexico, Central America, and northern and central ...
(''A. bicolor''), including by the
American Ornithological Society The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its m ...
and sometimes a distinct species. The two show differences in
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
preference and have
allopatric Allopatric speciation () – also referred to as geographic speciation, vicariant speciation, or its earlier name the dumbbell model – is a mode of speciation that occurs when biological populations become geographically isolated from ...
distributions. But the situation is complicated by the subspecies ''pileatus'', which is intermediate in plumage between ''bicolor'' and ''chilensis'', and has been variously assigned to either species by those that consider them distinct.


Description

The male is 37 to 38 cm long, while the larger female measures 41 to 42 cm. The adults have black upperparts, and an ash-grey chest and abdomen with dark barring. The throat has longitudinal dark stripes and the undertail is white. The uppertail is brown with 5 or 6 dark bands. The legs are greenish yellow, and the eyes are yellow. The sexes have similar plumage. Young birds have browner upperparts with cream fringes to the feathers. The paler chest and abdomen have longitudinal stripes. The paler uppertail makes the banding more obvious. Birds are only vocal during the breeding season. While engaging in reproductive activity, its calls do not seem to differ from those of the bicoloured hawk. This has at least two rather high-pitched scolding vocalizations: a barking row of ''keh'' or ''kow'', and a
woodpecker Woodpeckers are part of the bird family Picidae, which also includes the piculets, wrynecks, and sapsuckers. Members of this family are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar, and the extreme polar regions. ...
-like faster and accelerating
staccato Staccato (; Italian for "detached") is a form of musical articulation. In modern notation, it signifies a note of shortened duration, separated from the note that may follow by silence. It has been described by theorists and has appeared in music ...
of ''kek'' calls. These are given near the nest and might announce intruders such as the scientists coming to study the birds. A series of higher-pitched ''kie'', given by a definite Chilean hawk under similar circumstances, might replace one of the above, or it might be a distinct type of call. Pairmates address each other with a squealing ''waaah'', and a soft clear whistle is apparently used by parents to communicate something to the young.Trejo ''et al.'' (2006)


Distribution

In
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
, it is found north to
Neuquén Province Neuquén () is a province of Argentina, located in the west of the country, at the northern end of Patagonia. It borders Mendoza Province to the north, Rio Negro Province to the southeast, and Chile to the west. It also meets La Pampa Province a ...
at about 36°
latitude In geography, latitude is a coordinate that specifies the north– south position of a point on the surface of the Earth or another celestial body. Latitude is given as an angle that ranges from –90° at the south pole to 90° at the north pol ...
. The northern limit of the breeding range in
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
is not well known. In significant numbers, the Chilean hawk probably only occurs north to
O'Higgins Region The Libertador General Bernardo O'Higgins Region ( es, Región del Libertador General Bernardo O'Higgins, ), often shortened to O'Higgins Region ( es, Región de O'Higgins), is one of Chile's 16 first order administrative divisions. It is subdi ...
, though it at least occasionally breeds in
Valparaíso Region The Valparaíso Region ( es, Región de Valparaíso, links=no, ) is one of Chile's 16 first order administrative divisions.Valparaíso Region, 2006 With the country's second-highest population of 1,790,219 , and fourth-smallest area of , ...
, and perhaps also in
Coquimbo Region The Coquimbo Region ( es, Región de Coquimbo, ) is one of Chile's 16 regions (first order administrative divisions). It is bordered by Atacama to the north, Valparaíso to the south, Argentina to the east, and the Pacific Ocean to the west. ...
and Fray Jorge National Park from where there are a few sightings. The southern limit is generally accepted to be at about 55° latitude in
Tierra del Fuego Tierra del Fuego (, ; Spanish for "Land of the Fire", rarely also Fireland in English) is an archipelago off the southernmost tip of the South American mainland, across the Strait of Magellan. The archipelago consists of the main island, Isla G ...
. Their movement patterns are not well resolved. Generally it seems to be sedentary, but there is a seasonal influx of Chilean hawks to NW Argentina. Perhaps they follow the swarms of migrant
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by t ...
s or even
Chilean pigeon The Chilean pigeon (''Patagioenas araucana'') is a species of bird in the family Columbidae. It is found in Chile and Argentina. Taxonomy and systematics The Chilean pigeon is monotypic. It, ring-tailed pigeon (''Patagioenas caribaea''), and ...
s (''Columba araucana''). It has been recorded as breeding in
Magallanes Province Magallanes Province ( es, Provincia de Magallanes) is one of four Provinces of Chile, provinces in the southern Chilean Regions of Chile, region of Magallanes y la Antártica Chilena Region, Magallanes and Antártica Chilena. The provincial capital ...
but migrating away afterwards. Young birds seem to be quite prone to vagrancy: there is one certain and one possible record from
Tafí del Valle Tafí del Valle is a city in Tucumán, Argentina. It is located in the department of the same name, of which it is the main settlement, in the west of the Argentine province of Tucumán, 126 km from the provincial capital, San Miguel d ...
and
Oasis de Pica In ecology, an oasis (; ) is a fertile area of a desert or semi-desert environment'ksar''with its surrounding feeding source, the palm grove, within a relational and circulatory nomadic system.” The location of oases has been of critical im ...
, respectively; both locations are hundreds of kilometers north even of the suspected breeding range.


Ecology

The Chilean hawk is specifically found in
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
forest. Far more rarely, it is also found in
sclerophyll Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaf, leaves, short Internode (botany), internodes (the distance between leaves along the stem) and leaf orientation which is paral ...
ous forest, parkland and mixed forest and open habitat. Top hunt, it also visits open areas like shrubland, grassland or agricultural land to hunt. It is rarely if ever seen in heavily human-modified habitat however, and the few individuals that have been encountered in city parks and gardens are probably not resident birds. As it seems, it requires not much less than 200
hectare The hectare (; SI symbol: ha) is a non-SI metric unit of area equal to a square with 100-metre sides (1 hm2), or 10,000 m2, and is primarily used in the measurement of land. There are 100 hectares in one square kilometre. An acre is a ...
s of native forest to breed. Typically, forest inhabited by this bird is dominated by ''
Araucaria ''Araucaria'' (; original pronunciation: .ɾawˈka. ɾja is a genus of evergreen Conifer, coniferous trees in the family Araucariaceae. There are 20 extant taxon, extant species in New Caledonia (where 14 species are endemism, ende ...
'' and
southern beech ''Nothofagus'', also known as the southern beeches, is a genus of 43 species of trees and shrubs native to the Southern Hemisphere in southern South America (Chile, Argentina) and Australasia (east and southeast Australia, New Zealand, New Gui ...
(''Nothofagus''). Particular species that have been recorded are
coihue ''Nothofagus dombeyi'', Dombey's beech, coigue, coihue or coigüe (from Mapudungun ''koywe'') is a tree species native to southern Chile and the Andean parts of Argentine Patagonia. It is a fast-growing species that can live in a wide range of ...
(''N. dombeyi''), hualle (''N. obliqua'') and lenga (''N. pumilio''). It probably tolerates some
logging Logging is the process of cutting, processing, and moving trees to a location for transport. It may include skidding, on-site processing, and loading of trees or logs onto trucks or skeleton cars. Logging is the beginning of a supply chain ...
, as long as the native character of the forest is not altered.
Secondary growth In botany, secondary growth is the growth that results from cell division in the cambia or lateral meristems and that causes the stems and roots to thicken, while primary growth is growth that occurs as a result of cell division at the tips of ...
with abundant mature trees remaining and dense undergrowth, e.g. of South American mountain bamboo (''Chusquea''), as caused by limited logging activity, may even be prime breeding habitat, although too little is known to be certain. When enough native forest is present,
plantation A plantation is an agricultural estate, generally centered on a plantation house, meant for farming that specializes in cash crops, usually mainly planted with a single crop, with perhaps ancillary areas for vegetables for eating and so on. The ...
s, e.g. of introduced
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...
s, are also utilized. During the day, it likes to perch on branches in its territory, moving between favorite areas of forest in low flight. Areas with strong human activity like settlements are approached cautiously; it is generally not a bird that announces its presence. Pairs split outside the breeding season; it is not studied whether they are monogamous only during the breeding season or for several seasons. It seldom soars unrelated to reproductive activity. Males do aerobatic displays in courtship, such as a double loop resembling an upright "8". The
louse Louse ( : lice) is the common name for any member of the clade Phthiraptera, which contains nearly 5,000 species of wingless parasitic insects. Phthiraptera has variously been recognized as an order, infraorder, or a parvorder, as a result o ...
'' Colpocephalum turbinatum'' was found on a museum specimen of the Chilean hawk, but whether it actually parasitizes these birds or had simply crossed over from some other specimen is not known.


Food and feeding

It is not well known how this
carnivore A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
catches its prey, but as it seems it is optimized for pursuit of small and maneuvrable birds throughout all levels of the forest. It is also able to seize large
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s in mid-air. Both active searching for prey and sitting in ambush to wait what might come along has been observed. During the breeding season, pairs may cooperate in hunting; their different sizes ensures that they do not compete for prey much. The Chilean hawk's food is almost exclusively birds (97.8% of all prey remains in one study),Figueroa Rojas ''et al.'' (2004) in particular a diverse selection of forest
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by t ...
s. More than 30 bird species are documented to be eaten by this hawk at least occasionally.
Rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are na ...
s of at least 4 species and every now and then an occasional
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
or
squamate Squamata (, Latin ''squamatus'', 'scaly, having scales') is the largest order of reptiles, comprising lizards, snakes, and amphisbaenians (worm lizards), which are collectively known as squamates or scaled reptiles. With over 10,900 species, ...
round off its diet. The Chilean hawk hunts forest passerines quite indiscriminately of species, habitat or habits provided they have the right size, though it has a preference for species that live closer to the forest floor. Depending on availability, favorite prey species include thorn-tailed rayadito (''Aphrastura spinicauda'') black-chinned siskin (''Carduelis barbata''), white-crested elaenia (''Elaenia albiceps''),
Austral thrush The austral thrush (''Turdus falcklandii'') is a medium-sized thrush from southern South America. There are two subspecies, the Magellan thrush (''T. f. magellanicus'') from south Argentina and south and central Chile, and the Falkland thrush (' ...
(''Turdus falcklandii'') and fire-eyed diucon (''Xolmis pyrope''). It has been claimed that the
Chilean pigeon The Chilean pigeon (''Patagioenas araucana'') is a species of bird in the family Columbidae. It is found in Chile and Argentina. Taxonomy and systematics The Chilean pigeon is monotypic. It, ring-tailed pigeon (''Patagioenas caribaea''), and ...
(''Columba araucana'') constitutes important prey, but this seems only to be correct at certain times or places, if at all.


Reproduction

This hawk breeds in the
austral summer Summer is the hottest of the four temperate seasons, occurring after spring and before autumn. At or centred on the summer solstice, the earliest sunrise and latest sunset occurs, daylight hours are longest and dark hours are shortest, ...
. Pairs apparently form from mid-late October on. Incubating birds have been observed in December, and chicks are seen from about
New Year's Eve In the Gregorian calendar, New Year's Eve, also known as Old Year's Day or Saint Sylvester's Day in many countries, is the evening or the entire day of the last day of the year, on 31 December. The last day of the year is commonly referred to ...
to February, after which the families disperse again. A 1937 nest and a 1945 egg description of the Chilean hawk appear to be based on a misidentification; the nest at least was probably of the
chimango caracara The chimango caracara (''Milvago chimango'') is a species of bird of prey in the family Falconidae. It is found in Argentina, Uruguay, Chile, Paraguay and south of Brazil. The chimango is found as far south as Tierra del Fuego and is a vagrant t ...
(''Milvago chimango''). The oval platform nest measures about 50–80 by 50–60 cm and is some 25 cm high when freshly built. Some nests are more than twice as high; these might have been used in several years. It is built from strongly intertwined dry twigs and sticks. It is placed on forked branches in the upper part of a tree, close to the main trunk or a main vertical branch, some 16–20 m above ground. At least locally, full-grown
coihue ''Nothofagus dombeyi'', Dombey's beech, coigue, coihue or coigüe (from Mapudungun ''koywe'') is a tree species native to southern Chile and the Andean parts of Argentine Patagonia. It is a fast-growing species that can live in a wide range of ...
trees (''Nothofagus dombeyi'') seem to be much preferred for nesting. Nests are sometimes reused in successive seasons, but more often a new nest is constructed in a different tree every season.Trejo ''et al.'' (2006), Figueroa Rojas ''et al.'' (2007) The
clutch A clutch is a mechanical device that engages and disengages power transmission, especially from a drive shaft to a driven shaft. In the simplest application, clutches connect and disconnect two rotating shafts (drive shafts or line shafts). ...
is probably two, sometimes three and rarely one, as usual for
Accipitridae The Accipitridae is one of the three families within the order Accipitriformes, and is a family of small to large birds with strongly hooked bills and variable morphology based on diet. They feed on a range of prey items from insects to medium-s ...
. The eggs are dull light bluish to off-white all over and are shaped like a chicken's egg. The eggshell's inside has a slightly more pronounced bluish tinge. Incubation lasts probably about 3 weeks. The parents defend their nesting grounds against other birds of prey, such as the
red-backed hawk The variable hawk (''Geranoaetus polyosoma'') is a polymorphic species of bird of prey in the family Accipitridae. It is widespread and often common in open habitats in western and southern South America, including the Falkland Islands. Its ta ...
(''Buteo polyosoma''), and the
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird of p ...
(''Aquila chrysaetos''); during approaches by such potentially dangerous species, the nestlings will tuck away their heads. It seems that 2 or 3 young are raised on a regular basis, unlike in many other Accipitridae where only the strongest nestling survives.


Status

Because of its forest habitat and secretive behaviour, the Chilean hawk is one of the least-studied raptors in the
Patagonia Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and gl ...
n temperate forest. It is considered relatively common in the
Cape Horn Cape Horn ( es, Cabo de Hornos, ) is the southernmost headland of the Tierra del Fuego archipelago of southern Chile, and is located on the small Hornos Island. Although not the most southerly point of South America (which are the Diego Ramírez ...
region, e.g. in Ñuble National Reserve, population densities as high as 4 birds per square km have been recorded. Elsewhere, it is far less often seen and usually quite rare. Attempts to assess its
population density Population density (in agriculture: standing stock or plant density) is a measurement of population per unit land area. It is mostly applied to humans, but sometimes to other living organisms too. It is a key geographical term.Matt RosenberPopul ...
are hampered by the fact that it requires a certain amount of prime habitat to settle in a locale at all. Thus, much otherwise suitable land might be under-utilized by these birds, and
subpopulation In statistics, a population is a set of similar items or events which is of interest for some question or experiment. A statistical population can be a group of existing objects (e.g. the set of all stars within the Milky Way galaxy) or a hypothe ...
s have an extremely patchy distribution. It is listed as a rare or insufficiently known species in
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
and legally protected under the Hunting Law. In
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
it is not listed as threatened. On a global scale, it is a rare bird, though not under immediate threat. Populations may decline due to increasing habitat loss from extensive fires, logging pressures, and hunting. It is not evaluated by the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
, as they do not consider it specifically distinct, but is included on the
CITES CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of interna ...
Appendix II as part of the blanket listing of
Falconiformes The order Falconiformes () is represented by the extant family Falconidae (falcons and caracaras) and a handful of enigmatic Paleogene species. Traditionally, the other bird of prey families Cathartidae (New World vultures and condors), Sagitt ...
.


Footnotes


References

* Figueroa Rojas, Ricardo A.; Alvarado Orellana, Sergio; Corales Stappung, Soraya & Shehadeh, Ishback (2004): Prey of breeding Chilean hawks (''Accipiter chilensis'') in an Andean ''Nothofagus'' forest in northern Patagonia. '' Wilson Bull.'' 116(4): 347–351. (HTML abstract) * Figueroa Rojas, Ricardo A.; Alvarado Orellana, Sergio; Gonzalez-Acua, Daniel & Corales Stappung, Soraya (2007): Nest characteristics of the Chilean Hawk (''Accipiter chilensis'', Falconiformes: Accipitridae) in an Andean ''Nothofagus'' forest of northern Patagonia. ''Studies on Neotropical Fauna and Environment'' 42(1): 1-4 nglish with Spanish abstract (HTML abstract) * Trejo, Ana; Figueroa Rojas, Ricardo A. & Alvarado Orellana, Sergio (2006): Forest-specialist raptors of the temperate forests of southern South America: a review. ''Revista Brasileira de Ornitologia'' 14(4): 317-330 nglish with Portuguese abstractbr>PDF fulltext


External links


Accipiter Chilensis mating - Video by Emmy Award winner cinematographer Christian Munoz-Donoso
{{Taxonbar, from=Q114022
Chilean hawk The Chilean hawk (''Accipiter chilensis'') is a bird of prey species belonging to the typical hawks. It breeds in Andes forests from central Chile and western Argentina south to Tierra del Fuego, from sea level to 2,700 m altitude (though birds ...
Chilean hawk The Chilean hawk (''Accipiter chilensis'') is a bird of prey species belonging to the typical hawks. It breeds in Andes forests from central Chile and western Argentina south to Tierra del Fuego, from sea level to 2,700 m altitude (though birds ...
Birds of Chile Birds of the Southern Andes
Chilean hawk The Chilean hawk (''Accipiter chilensis'') is a bird of prey species belonging to the typical hawks. It breeds in Andes forests from central Chile and western Argentina south to Tierra del Fuego, from sea level to 2,700 m altitude (though birds ...
Chilean hawk The Chilean hawk (''Accipiter chilensis'') is a bird of prey species belonging to the typical hawks. It breeds in Andes forests from central Chile and western Argentina south to Tierra del Fuego, from sea level to 2,700 m altitude (though birds ...