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Chewing or mastication is the process by which
food Food is any substance consumed by an organism for nutritional support. Food is usually of plant, animal, or fungal origin, and contains essential nutrients, such as carbohydrates, fats, proteins, vitamins, or minerals. The substance is inge ...
is crushed and ground by
teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
. It is the first step of digestion, and it increases the surface area of foods to allow a more efficient break down by
enzyme Enzymes () are proteins that act as biological catalysts by accelerating chemical reactions. The molecules upon which enzymes may act are called substrates, and the enzyme converts the substrates into different molecules known as products. A ...
s. During the mastication process, the food is positioned by the cheek and
tongue The tongue is a muscular organ (anatomy), organ in the mouth of a typical tetrapod. It manipulates food for mastication and swallowing as part of the digestive system, digestive process, and is the primary organ of taste. The tongue's upper surfa ...
between the teeth for grinding. The
muscles of mastication There are four classical muscles of mastication. During mastication, three muscles of mastication (''musculi masticatorii'') are responsible for adduction of the jaw, and one (the lateral pterygoid) helps to abduct it. All four move the jaw late ...
move the
jaw The jaw is any opposable articulated structure at the entrance of the mouth, typically used for grasping and manipulating food. The term ''jaws'' is also broadly applied to the whole of the structures constituting the vault of the mouth and serv ...
s to bring the teeth into intermittent contact, repeatedly occluding and opening. As chewing continues, the food is made softer and warmer, and the enzymes in saliva begin to break down
carbohydrates In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may or may ...
in the food. After chewing, the food (now called a
bolus Bolus may refer to: Geography * Bolus, Iran, a village in Ardabil Province, Iran * Bolus, or Baulus, an Anatolian village on the site of ancient Berissa Medicine * Bolus (digestion), a ball-shaped mass moving through the digestive tract * Bolus ...
) is swallowed. It enters the
esophagus The esophagus (American English) or oesophagus (British English; both ), non-technically known also as the food pipe or gullet, is an organ in vertebrates through which food passes, aided by peristaltic contractions, from the pharynx to the ...
and via
peristalsis Peristalsis ( , ) is a radially symmetrical contraction and relaxation of muscles that propagate in a wave down a tube, in an anterograde direction. Peristalsis is progression of coordinated contraction of involuntary circular muscles, which ...
continues on to the stomach, where the next step of digestion occurs. Increasing the number of chews per bite increases relevant gut hormones. Studies suggest that chewing may decrease self-reported hunger and food intake. Chewing gum has been around for many centuries; there is evidence that northern Europeans chewed
birch bark tar Birch tar or birch pitch is a substance (liquid when heated) derived from the dry distillation of the bark of the birch tree. Compounds It is composed of phenols such as guaiacol, cresol, xylenol, and creosol. Ancient and modern uses B ...
9,000 years ago. Chewing, needing specialized teeth, is mostly a mammalian adaptation that appeared in early Synapsids, though some later herbivorous dinosaurs, since extinct, had developed chewing too. Nowadays, only mammals chew in the strict sense of the word, though some fishes have a somewhat similar behavior. Neither birds, nor amphibians or any living reptiles chew.
Premastication Premastication, pre-chewing, or kiss feeding is the act of chewing food for the purpose of physically breaking it down in order to feed another that is incapable of masticating the food by themselves. This is often done by the mother or relatives ...
is sometimes performed by human parents for infants who are unable to do so for themselves. The food is masticated in the mouth of the parent into a bolus and then transferred to the infant for consumption. (Some other animals also premasticate.)
Cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus ''Bos''. Adult females are referred to as cows and adult mal ...
and some other animals, called ruminants, chew food more than once to extract more nutrients. After the first round of chewing, this food is called
cud Cud is a portion of food that returns from a ruminant's stomach to the mouth to be chewed for the second time. More precisely, it is a bolus of semi-degraded food regurgitated from the reticulorumen of a ruminant. Cud is produced during the phy ...
. Chewing is important and beneficial for overall health. If food is not chewed correctly it can cause choking and other digestive problems.


Chewing motor program

Chewing is primarily an unconscious ( semi-autonomic) act, but can be mediated by higher conscious input. The motor program for mastication is a hypothesized central nervous system function by which the complex patterns governing mastication are created and controlled. It is thought that feedback from proprioceptive nerves in teeth and the
temporomandibular In anatomy, the temporomandibular joints (TMJ) are the two joints connecting the jawbone to the skull. It is a bilateral Synovial joint, synovial articulation between the temporal bone of the skull above and the Human mandible, mandible below; ...
joints govern the creation of neural pathways, which in turn determine duration and force of individual muscle activation (and in some cases muscle fiber groups as in the masseter and temporalis). This motor program continuously adapts to changes in food type or occlusion. This adaptation is a learned skill that may sometimes require relearning to adapt to loss of teeth or to dental appliances such as dentures. It is thought that conscious mediation is important in the limitation of parafunctional habits as most commonly, the motor program can be excessively engaged during periods of sleep and times of stress. It is also theorized that excessive input to the motor program from myofascial pain or occlusal imbalance can contribute to parafunctional habits.


Muscles


Nutrition and health

Chewing stimulates
saliva Saliva (commonly referred to as spit) is an extracellular fluid produced and secreted by salivary glands in the mouth. In humans, saliva is around 99% water, plus electrolytes, mucus, white blood cells, epithelial cells (from which DNA can be ...
production and increases sensory perception of the food being eaten, controlling when the food is swallowed. Evidence from one study suggests that chewing almonds 25-40 times kept people fuller while also allowing them to get more nutrients out of the almonds. The researchers also suggest that this is likely to be the case in other foods. Eating food which does not require chewing, by choice or due to medical reasons as tooth loss, is known as a
soft diet A mechanical soft diet or edentulous diet or soft food(s) diet is a diet that involves only foods that are physically soft, with the goal of reducing or eliminating the need to chew the food. It is recommended for people who have difficulty che ...
. Such a diet may lead to inadequate nutrition due to a reduction in fruit and vegetable intake. Chewing also stimulates the
hippocampus The hippocampus (via Latin from Greek , 'seahorse') is a major component of the brain of humans and other vertebrates. Humans and other mammals have two hippocampi, one in each side of the brain. The hippocampus is part of the limbic system, a ...
and is necessary to maintain its normal function.


In other animals

Chewing is largely an adaptation for
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
ian herbivory. Carnivores generally chew very little or swallow their food whole or in chunks. This act of gulping food (or medicine pills) without chewing has inspired the English
idiom An idiom is a phrase or expression that typically presents a figurative, non-literal meaning attached to the phrase; but some phrases become figurative idioms while retaining the literal meaning of the phrase. Categorized as formulaic language, ...
"wolfing it down". Other animals such as cows chew their food for long periods to allow for proper digestion in a process known as rumination. Rumination in cows has been shown by researchers to intensify during the night. They concluded that cows chewed more intently in the night time compared to the morning. Ornithopods, a group of
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s including the
Hadrosaurid Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which inclu ...
s ("duck-bills"), developed teeth analogous to mammalian molars and incisors during the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
period; this advanced, cow-like dentition allowed the creatures to obtain more nutrients from the tough plant life. This may have given them the advantage needed to compete with the formidable sauropods, who depended on their massive gastrointestinal tracts to digest food without grinding it, in their ecological niches. They eventually became some of the most successful animals on the planet until the
Cretaceous–Paleogene extinction event The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million years ago. With the ...
wiped them out.


In machinery

The process of chewing has, by analogy, been applied to machinery. The
U.S. Forest Service The United States Forest Service (USFS) is an agency of the U.S. Department of Agriculture that administers the nation's 154 national forests and 20 national grasslands. The Forest Service manages of land. Major divisions of the agency in ...
uses a machine called a ''masticator'' (also called a forestry mulching machine) to "chew" through brush and timber in order to clear
fireline A firebreak or double track (also called a fire line, fuel break, fireroad and firetrail in Australia) is a gap in vegetation or other combustible material that acts as a barrier to slow or stop the progress of a bushfire or wildfire. A firebre ...
s in advance of a wildfire. A cold press juicer uses the mastication process to extract juice from fruit and vegetable without the loss of oxygen or heat-sensitive nutrients as there is less friction involved.


See also

*
Biting Biting is a common zoological behavior involving the active, rapid closing of the jaw around an object. This behavior is found in toothed animals such as mammals, reptiles, amphibians and fish, but can also exist in arthropods. Myocytic contrac ...
* Gnathology *
Muscles of mastication There are four classical muscles of mastication. During mastication, three muscles of mastication (''musculi masticatorii'') are responsible for adduction of the jaw, and one (the lateral pterygoid) helps to abduct it. All four move the jaw late ...
* Horace Fletcher * Chewing Gum


Notes


External links

* {{Authority control Dentistry Digestive system Articles containing video clips sk:Žuvacie svaly