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A haemal arch also known as a chevron, is a bony arch on the ventral side of a tail
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristi ...
of a
vertebrate Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, ...
. The canal formed by the space between the arch and the vertebral body is the haemal canal. A spinous ventral process emerging from the haemal arch is referred to as the haemal spine. Blood vessels to and from the tail run through the arch. In reptiles, the caudofemoralis longus muscle, one of the main muscles involved in locomotion, attaches to the lateral sides of the haemal arches. In 1956,
Alfred Sherwood Romer Alfred Sherwood Romer (December 28, 1894 – November 5, 1973) was an American paleontologist and biologist and a specialist in vertebrate evolution. Biography Alfred Romer was born in White Plains, New York, the son of Harry Houston Romer an ...
hypothesized that the position of the first haemal arch was sexually dimorphic in crocodilians and dinosaurs. However, subsequent research established that the size and position of the first haemal arch was not sexually dimorphic in crocodilians and found no evidence of significant variation in tyrannosaurid dinosaurs, indicating that haemal arches could not be used to distinguish between sexes after all. Haemal arches play an important role in the taxonomy of sauropod dinosaurs, as sauropods exhibit a wide range of morphologies of the haemal arches. In 1878,
Othniel Marsh Othniel Charles Marsh (October 29, 1831 – March 18, 1899) was an American professor of Paleontology in Yale College and President of the National Academy of Sciences. He was one of the preeminent scientists in the field of paleontology. Among h ...
named the sauropod ''
Diplodocus ''Diplodocus'' (, , or ) was a genus of diplodocid sauropod dinosaurs, whose fossils were first discovered in 1877 by S. W. Williston. The generic name, coined by Othniel Charles Marsh in 1878, is a neo-Latin term derived from Greek Î´Î¹Ï ...
'' after the distinctive shape of its haemal arches, which were forked to have both an anterior and posterior process. Though once thought to be a specialized characteristic of ''Diplodocus'' and its close relatives, forked chevrons are now known to have been widespread among sauropod dinosaurs, although titanosauriform sauropods returned to the unforked condition. File:Wintonotitan tail chevrons.jpg, Haemal arches of ''
Wintonotitan ''Wintonotitan'' (meaning " Winton titan") is a genus of titanosauriform dinosaur from late Albian (Early Cretaceous)-age Winton Formation of Australia. It is known from partial postcranial remains. Description and history Fossils that are ...
''. File:Ichthyovenator laosensis tail vertebrae NMNS.jpg, Caudal vertebrae of ''
Ichthyovenator ''Ichthyovenator'' is a genus of spinosaurid dinosaur that lived in what is now Laos, sometime between 125 and 113 million years ago, during the Aptian stage of the Early Cretaceous period. It is known from fossils collected from the Grè ...
'', showing haemal arches below tail. File:The Childrens Museum of Indianapolis - Chevron from Edmontosaurus - detail.jpg, Haemal arch of '' Edmontosaurus''. File:Oksoko & Rinchenia tail chevrons.jpg, Haemal arch series of ''
Oksoko The three-headed eagle (or triple-headed eagle) is a mythological or heraldic bird, as it were an augmented version of the double-headed eagle. A three-headed eagle is mentioned in the apocryphal Latin Ezra, featuring in a dream by the high ...
'' and ''
Rinchenia ''Rinchenia'' (named after Byambyn Rinchen) is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous epoch in what is now Mongolia, Nemegt Formation, around 70 million years ago. The type and only known species, ''R ...
''. File:Ischioceratops caudal vertebrae.png, Caudal vertebrae of ''
Ischioceratops ''Ischioceratops'' () is an extinct genus of small herbivorous ceratopsian dinosaur that lived approximately 69 million years ago during the latter part of the Cretaceous Period in what is now China. ''Ischioceratops'' was a small sized, mode ...
'', showing haemal arches below tail.


References

* Vertebrate anatomy {{Vertebrate anatomy-stub