Cheirogaleid
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The Cheirogaleidae are the family of
strepsirrhine Strepsirrhini or Strepsirhini (; ) is a suborder of primates that includes the lemuriform primates, which consist of the lemurs of Madagascar, galagos ("bushbabies") and pottos from Africa, and the lorises from India and southeast Asia. Collec ...
primate Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians (monkeys and apes, the latter including huma ...
s containing the various dwarf and mouse lemurs. Like all other
lemur Lemurs ( ) (from Latin ''lemures'' – ghosts or spirits) are Strepsirrhini, wet-nosed primates of the Superfamily (biology), superfamily Lemuroidea (), divided into 8 Family (biology), families and consisting of 15 genera and around 100 exist ...
s, cheirogaleids live exclusively on the island of
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
.


Characteristics

Cheirogaleids are smaller than the other lemurs and, in fact, they are the smallest primates. They have soft, long fur, colored grey-brown to reddish on top, with a generally brighter underbelly. Typically, they have small ears, large, close-set eyes, and long hind legs. Like all strepsirrhines, they have fine claws at the second toe of the hind legs. They grow to a size of only 13 to 28 cm, with a tail that is very long, sometimes up to one and a half times as long as the body. They weigh no more than 500 grams, with some species weighing as little as 60 grams. Dwarf and mouse lemurs are
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
and
arboreal Arboreal locomotion is the Animal locomotion, locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. Th ...
. They are excellent climbers and can also jump far, using their long tails for balance. When on the ground (a rare occurrence), they move by hopping on their hind legs. They spend the day in tree hollows or leaf nests. Cheirogaleids are typically solitary, but sometimes live together in pairs. Their eyes possess a
tapetum lucidum The ''tapetum lucidum'' ( ; ; ) is a layer of tissue in the eye of many vertebrates and some other animals. Lying immediately behind the retina, it is a retroreflector. It reflects visible light back through the retina, increasing the light a ...
, a light-reflecting layer that improves their night vision. Some species, such as the lesser dwarf lemur, store fat at the hind legs and the base of the tail, and hibernate. Unlike lemurids, they have long upper
incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, whe ...
s, although they do have the comb-like teeth typical of all strepsirhines. They have the
dental formula Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiolo ...
: Cheirogaleids are
omnivore An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nutr ...
s, eating fruits, flowers and leaves (and sometimes nectar), as well as insects, spiders, and small vertebrates. The females usually have three pairs of nipples. After a meager 60-day gestation, they will bear two to four (usually two or three) young. After five to six weeks, the young are weaned and become fully mature near the end of their first year or sometime in their second year, depending on the species. In human care, they can live for up to 15 years, although their life expectancy in the wild is probably significantly shorter.


Classification

The five
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
of cheirogaleids contain 42
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
. * Infraorder Lemuriformes ** Family Cheirogaleidae *** Genus '' Cheirogaleus'': dwarf lemurs ****
Montagne d'Ambre dwarf lemur The Montagne d'Ambre dwarf lemur or Andy Sabin's dwarf lemur (''Cheirogaleus andysabini'') is a species of dwarf lemur known only from Amber Mountain National Park and the surrounding area in northern Madagascar. It was identified in 2005, but n ...
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Cheirogaleus andysabini The Montagne d'Ambre dwarf lemur or Andy Sabin's dwarf lemur (''Cheirogaleus andysabini'') is a species of dwarf lemur known only from Amber Mountain National Park and the surrounding area in northern Madagascar. It was identified in 2005, but n ...
'' ****
Furry-eared dwarf lemur The furry-eared dwarf lemur (''Cheirogaleus crossleyi''), or Crossley's dwarf lemur, only found on the island of Madagascar, as with all other lemurs. It has a pelage Fur is a thick growth of hair that covers the skin of mammals. It consists ...
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Cheirogaleus crossleyi The furry-eared dwarf lemur (''Cheirogaleus crossleyi''), or Crossley's dwarf lemur, only found on the island of Madagascar, as with all other lemurs. It has a pelage Fur is a thick growth of hair that covers the skin of mammals. It consists ...
'' ****
Groves' dwarf lemur Groves' dwarf lemur (''Cheirogaleus grovesi'') is a species of dwarf lemur known from Ranomafana and Andringitra national parks and surrounding areas in southeastern Madagascar. The identification was based on comparison of mitochondrial and n ...
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Cheirogaleus grovesi Groves' dwarf lemur (''Cheirogaleus grovesi'') is a species of dwarf lemur known from Ranomafana and Andringitra national parks and surrounding areas in southeastern Madagascar. The identification was based on comparison of mitochondrial and n ...
'' ****
Lavasoa dwarf lemur The Lavasoa dwarf lemur (''Cheirogaleus lavasoensis'') is a small, nocturnal strepsirrhine primate and a species of lemur that is endemic to three small, isolated patches of forest on the southern slopes of the Lavasoa Mountains in southern Madaga ...
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Cheirogaleus lavasoensis The Lavasoa dwarf lemur (''Cheirogaleus lavasoensis'') is a small, nocturnal strepsirrhine primate and a species of lemur that is endemic to three small, isolated patches of forest on the southern slopes of the Lavasoa Mountains in southern Madaga ...
'' ****
Greater dwarf lemur The greater dwarf lemur (''Cheirogaleus major''), or the Geoffroy's dwarf lemur, is a lemur that is widely distributed over the primary and secondary forests near the eastern coast of Madagascar. They are also found in northern parts of Madagasca ...
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Cheirogaleus major The greater dwarf lemur (''Cheirogaleus major''), or the Geoffroy's dwarf lemur, is a lemur that is widely distributed over the primary and secondary forests near the eastern coast of Madagascar. They are also found in northern parts of Madagasca ...
'' ****
Fat-tailed dwarf lemur The fat-tailed dwarf lemur (''Cheirogaleus medius''), also known as the lesser dwarf lemur, western fat-tailed dwarf lemur, or spiny forest dwarf lemur, is endemic to Madagascar. Hibernation Recent research has shown that ''C. medius'' hibernat ...
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Cheirogaleus medius The fat-tailed dwarf lemur (''Cheirogaleus medius''), also known as the lesser dwarf lemur, western fat-tailed dwarf lemur, or spiny forest dwarf lemur, is endemic to Madagascar. Hibernation Recent research has shown that ''C. medius'' hibernat ...
'' ****
Lesser iron-gray dwarf lemur The lesser iron-gray dwarf lemur (''Cheirogaleus minusculus''), or large iron-gray dwarf lemur, is a small nocturnal lemur endemic to Madagascar. It is nocturnal and an arboreal quadruped, and is not very agile. It produces three types of vocalisa ...
, '' Cheirogaleus minusculus'' ****
Ankarana dwarf lemur The Ankarana dwarf lemur or Sheth's dwarf lemur (''Cheirogaleus shethi'') is a species of dwarf lemur, part of the ''C. medius'' group, known only from northern Madagascar. It has been found in two reserves (Ankarana Special Reserve and Analamera ...
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Cheirogaleus shethi The Ankarana dwarf lemur or Sheth's dwarf lemur (''Cheirogaleus shethi'') is a species of dwarf lemur, part of the ''C. medius'' group, known only from northern Madagascar. It has been found in two reserves (Ankarana Special Reserve and Analamera ...
'' ****
Sibree's dwarf lemur Sibree's dwarf lemur (''Cheirogaleus sibreei'') is a small nocturnal lemur endemic to Madagascar. The name of this dwarf lemur commemorates the English missionary and naturalist James Sibree (1836–1929). In 2010, a research team confirmed tha ...
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Cheirogaleus sibreei Sibree's dwarf lemur (''Cheirogaleus sibreei'') is a small nocturnal lemur endemic to Madagascar. The name of this dwarf lemur commemorates the English missionary and naturalist James Sibree (1836–1929). In 2010, a research team confirmed tha ...
'' ****
Thomas' dwarf lemur The Thomas' dwarf lemur (''Cheirogaleus thomasi'') is a species of dwarf lemur known only from Anosy, Madagascar. It lives in forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout th ...
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Cheirogaleus thomasi The Thomas' dwarf lemur (''Cheirogaleus thomasi'') is a species of dwarf lemur known only from Anosy, Madagascar. It lives in forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the ...
'' *** Genus ''
Microcebus The mouse lemurs are nocturnal lemurs of the genus ''Microcebus''. Like all lemurs, mouse lemurs are native to Madagascar. Mouse lemurs have a combined head, body and tail length of less than , making them the smallest primates (the smallest spe ...
'': mouse lemurs ****
Arnhold's mouse lemur Arnhold's mouse lemur or the Montagne d'Ambre mouse lemur (''Microcebus arnholdi'') is a species of mouse lemur endemic to Madagascar. Its holotype was first collected on 27 November 2005, and was first described in 2008. According to genetic t ...
, '' Microcebus arnholdi'' ****
Madame Berthe's mouse lemur Madame Berthe's mouse lemur (''Microcebus berthae'') or Berthe's mouse lemur is the smallest of the mouse lemurs and the smallest primate in the world; the average body length is and seasonal weight is around . ''Microcebus berthae'' is one of ...
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Microcebus berthae Madame Berthe's mouse lemur (''Microcebus berthae'') or Berthe's mouse lemur is the smallest of the mouse lemurs and the smallest primate in the world; the average body length is and seasonal weight is around . ''Microcebus berthae'' is one of ...
'' ****
Bongolava mouse lemur The Bongolava mouse lemur (''Microcebus bongolavensis'') is a species of mouse lemur endemic to Madagascar. It lives in western deciduous forest within a limited range, including Bongolava Forest and Ambodimahabibo Forest between the Sofia Riv ...
'' Microcebus bongolavensis'' ****
Boraha mouse lemur The Nosy Boraha mouse lemur (''Microcebus boraha'') is a species of mouse lemur described in 2016 from Madagascar. It was discovered by a team of researchers at the German Primate Center. It was initially discovered among closely related species ...
'' Microcebus boraha'' ****
Danfoss' mouse lemur Danfoss's mouse lemur (''Microcebus danfossi''), or the Ambarijeby mouse lemur, is a species of mouse lemur endemic to Madagascar. It lives in western deciduous forest within a limited range between the Sofia River and the Maevarano River. This ...
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Microcebus danfossi Danfoss's mouse lemur (''Microcebus danfossi''), or the Ambarijeby mouse lemur, is a species of mouse lemur endemic to Madagascar. It lives in western deciduous forest within a limited range between the Sofia River and the Maevarano River. This ...
'' ****
Ganzhorn's mouse lemur Ganzhorn's mouse lemur (''Microcebus ganzhorni'') is a species of lemur described in 2016. The discovery was made by researchers at the German Primate Center. It was discovered in Madagascar among closely related species such as Madame Berthe's m ...
. '' Microcebus ganzhorni'' ****
Gerp's mouse lemur Gerp's mouse lemur (''Microcebus gerpi'') is a species of mouse lemur known only from the Sahafina Forest in eastern Madagascar, near Mantadia National Park. Its discovery was announced in 2012 by a German and Malagasy research team. The Saha ...
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Microcebus gerpi Gerp's mouse lemur (''Microcebus gerpi'') is a species of mouse lemur known only from the Sahafina Forest in eastern Madagascar, near Mantadia National Park. Its discovery was announced in 2012 by a German and Malagasy research team. The Saha ...
'' ****
Reddish-gray mouse lemur The reddish-gray mouse lemur (''Microcebus griseorufus'') also known as the gray-brown mouse lemur or rufous-gray mouse lemur, is a small, solitary-but-social, nocturnal and omnivorous primate from the Western Madagascar in the region around Bez ...
, '' Microcebus griseorufus'' ****
Jolly's mouse lemur Jolly's mouse lemur (''Microcebus jollyae'') is a species of mouse lemur from Mananjary and Kianjavato in Madagascar. The species is named in honor of primatologist Alison Jolly. Jolly's mouse lemur is uniformly reddish-brown with a small white ...
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Microcebus jollyae Jolly's mouse lemur (''Microcebus jollyae'') is a species of mouse lemur from Mananjary and Kianjavato in Madagascar. The species is named in honor of primatologist Alison Jolly. Jolly's mouse lemur is uniformly reddish-brown with a small white ...
'' ****
Jonah's mouse lemur ''Microcebus jonahi'', or Jonah's mouse lemur, is tiny species of primate. It weighs 60 g and has a body length of around 13 cm and its tail measures around 13 cm as well. It is the 25th recognized species of mouse lemur and the 1 ...
, '' Microcebus jonahi'' ****
Goodman's mouse lemur Goodman's mouse lemur (''Microcebus lehilahytsara'') is a species of mouse lemur from the region near Andasibe in eastern Madagascar. The species is named in honor of primatologist Steven M. Goodman. "Lehilahytsara" is a combination of the Ma ...
, '' Microcebus lehilahytsara'' ****
MacArthur's mouse lemur MacArthur's mouse lemur (''Microcebus macarthurii''), or the Anjiahely mouse lemur, is a species of mouse lemur known only from Makira Natural Park in northeastern Madagascar. The species is named in honor of the founder of the MacArthur Foundat ...
, '' Microcebus macarthurii'' ****
Claire's mouse lemur Claire's mouse lemur (''Microcebus mamiratra''), or the Nosy Be mouse lemur, is a newly described species of lemur from the genus of the mouse lemurs (''Microcebus''). It lives on the island Nosy Bé in the Antsiranana province of Madagascar, an ...
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Microcebus mamiratra Claire's mouse lemur (''Microcebus mamiratra''), or the Nosy Be mouse lemur, is a newly described species of lemur from the genus of the mouse lemurs (''Microcebus''). It lives on the island Nosy Bé in the Antsiranana province of Madagascar, an ...
'', synonymous to ''M. lokobensis'' ****
Bemanasy mouse lemur The Bemanasy mouse lemur, ''Microcebus manitatra'', is a species of mouse lemur described in 2016 from Madagascar. It was discovered by a team of researchers at the German Primate Center. It was initially discovered among closely related species ...
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Microcebus manitatra The Bemanasy mouse lemur, ''Microcebus manitatra'', is a species of mouse lemur described in 2016 from Madagascar. It was discovered by a team of researchers at the German Primate Center. It was initially discovered among closely related species ...
'' ****
Margot Marsh's mouse lemur Margot Marsh's mouse lemur or the Antafondro mouse lemur (''Microcebus margotmarshae'') is a species of mouse lemur endemic to Madagascar. Its holotype was first collected on 21 May 2006, proposed in 2006 by Andriantompohavana ''et al.'', and wa ...
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Microcebus margotmarshae Margot Marsh's mouse lemur or the Antafondro mouse lemur (''Microcebus margotmarshae'') is a species of mouse lemur endemic to Madagascar. Its holotype was first collected on 21 May 2006, proposed in 2006 by Andriantompohavana ''et al.'', and wa ...
'' ****
Marohita mouse lemur The Marohita mouse lemur (''Microcebus marohita'') is a species of mouse lemur known only from the Marohita Forest in eastern Madagascar, near the village of Marolambo. Specimens were first collected in December 2003, and its discovery was anno ...
, '' Microcebus marohita'' ****
Mittermeier's mouse lemur Mittermeier's mouse lemur (''Microcebus mittermeieri'') is a species of mouse lemur known only from Anjanaharibe-Sud Special Reserve in Madagascar. Mittermeier's mouse lemur is one of the smallest mouse lemurs, but is larger than Madame Berthe's ...
, '' Microcebus mittermeieri'' ****
Gray mouse lemur The gray mouse lemur (''Microcebus murinus''), grey mouse lemur or lesser mouse lemur, is a small lemur, a type of strepsirrhine primate, found only on the island of Madagascar. Weighing , it is the largest of the mouse lemurs (genus ''Microceb ...
, ''
Microcebus murinus The gray mouse lemur (''Microcebus murinus''), grey mouse lemur or lesser mouse lemur, is a small lemur, a type of strepsirrhine primate, found only on the island of Madagascar. Weighing , it is the largest of the mouse lemurs (genus ''Microceb ...
'' ****
Pygmy mouse lemur The pygmy mouse lemur (''Microcebus myoxinus''), also known as Peters' mouse lemur or dormouse lemur, is a primate weighing only ; it is the second smallest of the mouse lemurs. Its dorsal side is a rufous-brown colour, and creamy-white ventrally ...
, '' Microcebus myoxinus'' ****
Golden-brown mouse lemur The golden-brown mouse lemur (''Microcebus ravelobensis''), also known as the (Lac) Ravelobe mouse lemur, is part of the Cheirogaleidae family, and the smallest species of lemur. It is arboreal, nocturnal and usually social.Myers, P"Cheirogalei ...
, ''
Microcebus ravelobensis The golden-brown mouse lemur (''Microcebus ravelobensis''), also known as the (Lac) Ravelobe mouse lemur, is part of the Cheirogaleidae family, and the smallest species of lemur. It is arboreal, nocturnal and usually social.Myers, P"Cheirogalei ...
'' ****
Brown mouse lemur The brown mouse lemur (''Microcebus rufus'') is a small primate, and like the other mouse lemurs can only be found on the island of Madagascar. They are known also as the rufous mouse lemur, eastern rufous mouse lemur, red mouse lemur, or russet ...
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Microcebus rufus The brown mouse lemur (''Microcebus rufus'') is a small primate, and like the other mouse lemurs can only be found on the island of Madagascar. They are known also as the rufous mouse lemur, eastern rufous mouse lemur, red mouse lemur, or russet ...
'' ****
Sambirano mouse lemur The Sambirano mouse lemur (''Microcebus sambiranensis'') is a small, recently discovered primate and like the other mouse lemurs can only be found on the island of Madagascar. The dorsal side is both cinnamon and rufous-cinnamon and is grey ventr ...
, '' Microcebus sambiranensis'' ****
Simmons' mouse lemur Simmons' mouse lemur (''Microcebus simmonsi'') is a species of mouse lemur known only from Betampona Special Reserve, Analalava Special Community Reserve near Foulpointe, and Zahamena National Park in Madagascar. The species is named in honor of ...
, '' Microcebus simmonsi'' ****
Anosy mouse lemur The Anosy mouse lemur (''Microcebus tanosi'') is a species of mouse lemur known only from the Manantantely Forest and Ivorona Forest in southeastern Madagascar, near Tôlanaro. Specimens were first collected in April 2007, and its discovery wa ...
. '' Microcebus tanosi'' ****
Northern rufous mouse lemur The northern rufous mouse lemur (''Microcebus tavaratra''), northern brown mouse lemur, or Tavaratra mouse lemur is found in northern Madagascar from the Ankarana Special Reserve in the west to the Manambato River in the northeast, and up to the ...
, ''
Microcebus tavaratra The northern rufous mouse lemur (''Microcebus tavaratra''), northern brown mouse lemur, or Tavaratra mouse lemur is found in northern Madagascar from the Ankarana Special Reserve in the west to the Manambato River in the northeast, and up to the ...
'' *** Genus ''
Mirza Mirza may refer to: * Mirza, Kamrup, town in Assam, India * Mirza (name), historical royal title & noble * ''Mirza'', the genus of giant mouse lemur * "Mirza", song by Nino Ferrer * ''Mirza – The Untold Story'', Punjabi action romance film wri ...
'': giant mouse lemurs ****
Coquerel's giant mouse lemur Coquerel's giant mouse lemur (''Mirza coquereli''), also known as Coquerel's dwarf lemur or the southern giant mouse lemur, is a small nocturnal lemur endemic to Madagascar. This species can be found in parts of the Madagascar dry deciduous fore ...
or Coquerel's dwarf lemur, ''
Mirza coquereli Coquerel's giant mouse lemur (''Mirza coquereli''), also known as Coquerel's dwarf lemur or the southern giant mouse lemur, is a small nocturnal lemur endemic to Madagascar. This species can be found in parts of the Madagascar dry deciduous fores ...
'' ****
Northern giant mouse lemur The northern giant mouse lemur (''Mirza zaza''), or northern dwarf lemur, is a species of lemur discovered in 2005.Mirza zaza'' *** Genus '' Allocebus'' ****
Hairy-eared dwarf lemur The hairy-eared dwarf lemur (''Allocebus trichotis''), or hairy-eared mouse lemur, is one of the most scarcely known lemurs. ''A. trichotis'' is a nocturnal lemur that is endemic to Madagascar. It was originally named by Albert Günther in 1875 ...
, ''
Allocebus trichotis The hairy-eared dwarf lemur (''Allocebus trichotis''), or hairy-eared mouse lemur, is one of the most scarcely known lemurs. ''A. trichotis'' is a nocturnal lemur that is endemic to Madagascar. It was originally named by Albert Günther in 1875 a ...
'' *** Genus '' Phaner'': fork-marked lemurs ****
Masoala fork-marked lemur The Masoala fork-marked lemur (''Phaner furcifer''), also known as the eastern fork-marked lemur or Masoala fork-crowned lemur, is a species of lemur found in the coastal forests of northeastern Madagascar. It is a small nocturnal animal with lar ...
, '' Phaner furcifer'' ****
Pale fork-marked lemur The pale fork-marked lemur (''Phaner pallescens''), or western fork-marked lemur, is a species of lemur known from western Madagascar; south of the Fiherenana River to the region of Soalala. It is listed on CITES Appendix I. References

...
, ''
Phaner pallescens The pale fork-marked lemur (''Phaner pallescens''), or western fork-marked lemur, is a species of lemur known from western Madagascar; south of the Fiherenana River to the region of Soalala. It is listed on CITES CITES (shorter name for the ...
'' **** Pariente's fork-marked lemur, '' Phaner parienti'' ****
Amber Mountain fork-marked lemur The Amber Mountain fork-marked lemur (''Phaner electromontis''), also known as the Montagne d'Ambre fork-crowned lemur or Tanta, is a small primate, and like the other lemurs, can only be found on the island of Madagascar. The species is named aft ...
, ''
Phaner electromontis The Amber Mountain fork-marked lemur (''Phaner electromontis''), also known as the Montagne d'Ambre fork-crowned lemur or Tanta, is a small primate, and like the other lemurs, can only be found on the island of Madagascar. The species is named aft ...
''


Footnotes

*According to the letter of the
International Code of Zoological Nomenclature The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its publisher, the ...
, the correct name for this family should be Microcebidae, but the name Cheirogaleidae has been retained for stability. *In 2008, 7 new species of ''Microcebus'' were formally recognized, but ''Microcebus lokobensis'' ( Lokobe mouse lemur) was not among the additions, even though it was described in 2006. Therefore, its status as a species is still questionable.


References

{{Taxonbar, from=Q271785 Lemurs Primate families Taxa named by John Edward Gray Taxa described in 1873