Charaxes Richelmanni
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__NOTOC__ ''Charaxes richelmanni'' is a
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
in the family
Nymphalidae The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to the superfamily Papilionoidea, they are usually medium-sized to large butterflies. Most species have a red ...
. It is found in
Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...
,
Gabon Gabon (; ; snq, Ngabu), officially the Gabonese Republic (french: République gabonaise), is a country on the west coast of Central Africa. Located on the equator, it is bordered by Equatorial Guinea to the northwest, Cameroon to the north ...
, the
Republic of Congo The Republic of the Congo (french: République du Congo, ln, Republíki ya Kongó), also known as Congo-Brazzaville, the Congo Republic or simply either Congo or the Congo, is a country located in the western coast of Central Africa to the w ...
, the
Central African Republic The Central African Republic (CAR; ; , RCA; , or , ) is a landlocked country in Central Africa. It is bordered by Chad to the north, Sudan to the northeast, South Sudan to the southeast, the DR Congo to the south, the Republic of th ...
, the
Democratic Republic of Congo The Democratic Republic of the Congo (french: République démocratique du Congo (RDC), colloquially "La RDC" ), informally Congo-Kinshasa, DR Congo, the DRC, the DROC, or the Congo, and formerly and also colloquially Zaire, is a country in ...
and
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...
.Afrotropical Butterflies: File H - Charaxinae - Tribe Charaxini
/ref> The habitat consists of lowland evergreen forests.


Description

Van Someren. 1970 describes and figures (plate 7) ''C. richelmanni''


Subspecies

*''C. r. richelmanni'' (southern Cameroon, Gabon, Congo, Central African Republic, Democratic Republic of Congo) *''C. r. scheveni'' Ackery, 1995.(Tanzania) '' C. (richelmanni) ducarmei'' Plantrou, 1982Plantrou, J. 1982. Description de 9 sous-especes du genre ''Charaxes'' recemment decouvertes en Afrique (Lep. Nymphalidae). ''Miscellanea Entomologica'' 49: 101-115. (Democratic Republic of Congo: Kivu) is now considered as bona species. tat.Rev.2005/small>


Related species

Historical attempts to assemble a cluster of presumably related species into a "''Charaxes jasius'' Group" have not been wholly convincing. More recent taxonomic revision, corroborated by phylogenetic research, allow a more rational grouping congruent with cladistic relationships. Within a well-populated
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
of 27 related species sharing a common ancestor approximately 16 mya during the
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
, 26 are now considered together as The ''jasius'' Group. One of the two lineages within this clade forms a robust
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
group of seven species sharing a common ancestor approximately 2-3 mya, i.e. during the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58"Out of Africa again: A phylogenetic hypothesis of the genus ''Charaxes (Lepidoptera: Nymphalidae)'' based on five gene regions"
Aduse-Poku, Vingerhoedt, Wahlberg. Molecular Phylogenetics and Evolution (2009) 53;463–478
and are considered as the ''jasius'' subgroup. The second lineage leads to 19 other species within the Jasius group, which are split in to three well-populated subgroups of closely related species. The ''jasius'' Group (26 Species): Clade 1: ''jasius'' subgroup (7 species) Clade 2: contains the well-populated three additional subgroups (19 species) of the jasius Group: called the ''brutus'', ''pollux'', and ''eudoxus'' subgroups. *the ''eudoxus'' subgroup (11 species): *'' Charaxes eudoxus'' *'' Charaxes lucyae'' *''Charaxes richelmanni'' *'' Charaxes musakensis'' *'' Charaxes biokensis'' tat.rev.2005ref name="Turlin 2005"/> *'' Charaxes ducarmei'' *'' Charaxes druceanus'' *'' Charaxes tectonis'' *''
Charaxes phraortes ''Charaxes phraortes'' is a butterfly in the family Nymphalidae. It is found in north-eastern Madagascar. The habitat consists of wet coastal forests. Description Similar to '' pollux'' and ''phoebus'', but has the tails on the hindwing much lo ...
'' *''
Charaxes andranodorus __NOTOC__ ''Charaxes andranodorus'' is a butterfly in the family Nymphalidae. It is found on Madagascar. Description This rare species is one of the most magnificent forms of ''Charaxes''. Above it agrees very nearly with ''phraortes'', the bas ...
'' *'' Charaxes andrefana'' tat.rev.2025ref name="Turlin 2005"/> Further exploration of the phylogenetic relationships amongst existing ''Charaxes'' taxa is required to improve clarity.


References

*
Victor Gurney Logan Van Someren Victor Gurney Logan Van Someren (1886 in Melbourne – 24 July 1976) was a zoologist and entomologist. Van Someren was born in Australia. He attended George Watson's College and studied zoology at University of Edinburgh. He was also a dentist ...
, 1970 Revisional notes on African ''Charaxes'' (Lepidoptera: Nymphalidae). Part VI. ''Bulletin of the British Museum'' (Natural History) (Entomology)197-25


External links


Images of ''C. richelmanni ducarmei''
Royal Museum for Central Africa The Royal Museum for Central Africa or RMCA ( nl, Koninklijk Museum voor Midden-Afrika or KMMA; french: Musée royal de l'Afrique centrale or MRAC; german: Königliches Museum für Zentralafrika or KMZA), also officially known as the AfricaMuse ...
(Albertine Rift Project)
Images of ''C. richelmanni richelmanni''
(Albertine Rift Project)
''Charaxes richelmanni'' images
at
Consortium for the Barcode of Life The Consortium for the Barcode of Life (CBOL) was an international initiative dedicated to supporting the development of DNA barcoding as a global standard for species identification. CBOL's Secretariat Office is hosted by the National Museum of ...
includes verso
''Charaxes ducarmei'' images
at Consortium for the Barcode of Life Butterflies described in 1936 richelmanni {{Charaxinae-stub