''Charaxes pleione'', the common orange charaxes, or square-winged red charaxes, is a

butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...

in the family

Nymphalidae The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to the superfamily Papilionoidea, they are usually medium-sized to large butterflies. Most species have a red ...

. It is found in

Guinea Guinea ( ),, fuf, 𞤘𞤭𞤲𞤫, italic=no, Gine, wo, Gine, nqo, ߖߌ߬ߣߍ߫, bm, Gine officially the Republic of Guinea (french: République de Guinée), is a coastal country in West Africa. It borders the Atlantic Ocean to the we ...

Sierra Leone Sierra Leone,)]. officially the Republic of Sierra Leone, is a country on the southwest coast of West Africa. It is bordered by Liberia to the southeast and Guinea surrounds the northern half of the nation. Covering a total area of , Sierra ...

Liberia Liberia (), officially the Republic of Liberia, is a country on the West African coast. It is bordered by Sierra Leone to Liberia–Sierra Leone border, its northwest, Guinea to its north, Ivory Coast to its east, and the Atlantic Ocean ...

Ivory Coast Ivory Coast, also known as Côte d'Ivoire, officially the Republic of Côte d'Ivoire, is a country on the southern coast of West Africa. Its capital is Yamoussoukro, in the centre of the country, while its largest city and economic centre is ...

Ghana Ghana (; tw, Gaana, ee, Gana), officially the Republic of Ghana, is a country in West Africa. It abuts the Gulf of Guinea and the Atlantic Ocean to the south, sharing borders with Ivory Coast in the west, Burkina Faso in the north, and To ...

Nigeria Nigeria ( ), , ig, Naìjíríyà, yo, Nàìjíríà, pcm, Naijá , ff, Naajeeriya, kcg, Naijeriya officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf o ...

Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...

Gabon Gabon (; ; snq, Ngabu), officially the Gabonese Republic (french: République gabonaise), is a country on the west coast of Central Africa. Located on the equator, it is bordered by Equatorial Guinea to the northwest, Cameroon to the north ...

, the

Republic of the Congo The Republic of the Congo (french: République du Congo, ln, Republíki ya Kongó), also known as Congo-Brazzaville, the Congo Republic or simply either Congo or the Congo, is a country located in the western coast of Central Africa to the w ...

, the

Central African Republic The Central African Republic (CAR; ; , RCA; , or , ) is a landlocked country in Central Africa. It is bordered by Chad to the north, Sudan to the northeast, South Sudan to the southeast, the DR Congo to the south, the Republic of th ...

Angola , national_anthem = " Angola Avante"() , image_map = , map_caption = , capital = Luanda , religion = , religion_year = 2020 , religion_ref = , coordina ...

, the

Democratic Republic of the Congo The Democratic Republic of the Congo (french: République démocratique du Congo (RDC), colloquially "La RDC" ), informally Congo-Kinshasa, DR Congo, the DRC, the DROC, or the Congo, and formerly and also colloquially Zaire, is a country in ...

Uganda }), is a landlocked country in East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territor ...

Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...

Rwanda Rwanda (; rw, u Rwanda ), officially the Republic of Rwanda, is a landlocked country in the Great Rift Valley of Central Africa, where the African Great Lakes region and Southeast Africa converge. Located a few degrees south of the Equator ...

and

Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...

.Afrotropical Butterflies: File H - Charaxinae - Tribe Charaxini
/ref>

Description

Hindwing between the anal angle and vein 4 nearly straight, at vein 2 feebly angled but not tailed, at vein 4 with a short tail, only about 3 mm. in length. Wings above red-brown; apex and distal margin of the forewing more or less broadly black. Wings beneath with a common dark transverse line, extending from the costal margin of the forewing, just before the apex, to the anal angle of the hindwing. Costal margin of the forewing beneath in cellule 12 white or whitish to the end of the cell.-''lichas'' Dbl. male The black colour of the apical area of the forewing extends to the apex of the cell and is either spotted at the inner margin or quite unicolorous - ''bebra'' Rothsch. male The black colour at the apex of the forewing does not nearly reach the cell and is deeply incised or spotted with the ground-colour, especially in cellules 4 and 5. Cameroons, Congo region, German East Africa. Similar to '' Charaxes paphianus'' but forewing apex not attenuated.

Biology

The habitat consists of forests and woodland savanna. The larvae feed on ''

Acacia pennata ''Senegalia pennata'' ( en, climbing wattle, vi, rau thối, th, ชะอม ''cha-om'', my, ဆူးပုပ်, ; km, ស្អំ; Meiteilon : ''khang'', Thadou-Kuki: khang-khu, Paite Language: Khangkhuh, Mizo: khanghu, Hmar: ''khan ...

'' and '' Acacia brevispica''. Notes on the biology of ''pleione'' are given by Kielland (1990) Larsen (1991) and Larsen (2005)

Subspecies

*''C. p. pleione'' — Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, Nigeria *''C. p. bebra'' Rothschild, 1900 — north-eastern Democratic Republic of the Congo, Uganda, south-western Kenya, north-western Tanzania *''C. p. congoensis'' Plantrou, 1989 — south-eastern Nigeria, Cameroon, Gabon, Congo, Central African Republic, Angola, Democratic Republic of the Congo *''C. p. delvauxi'' Turlin, 1987 Turlin , B. 1987. Descriptions de nouvelles sous-especes et d’une forme de ''Charaxes'' africains (Lepidoptera Nymphalidae). ''Bulletin de la Societe Scientifique Naturelle'' No. 53: 19-25. — south-western Rwanda *''C. p. oriens'' Plantrou, 1989 — coast of Kenya, eastern Tanzania

Taxonomy

''Charaxes pleione'' is a member of the

species group In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...

'' Charaxes paphianus''. The supposed

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...

members are: *'' Charaxes paphianus'' - nominate *''Charaxes pleione''

Realm

Afrotropical realm The Afrotropical realm is one of Earth's eight biogeographic realms. It includes Africa south of the Sahara Desert, the majority of the Arabian Peninsula, the island of Madagascar, southern Iran and extreme southwestern Pakistan, and the island ...

References

Victor Gurney Logan Van Someren Victor Gurney Logan Van Someren (1886 in Melbourne – 24 July 1976) was a zoologist and entomologist. Van Someren was born in Australia. He attended George Watson's College and studied zoology at University of Edinburgh. He was also a dentist ...

, Revisional notes on African ''Charaxes'' (Lepidoptera: Nymphalidae). Part IX. ''Bulletin of the British Museum of Natural History'' (Entomology) 29 (8):415-487
Seitz, A. ''Die Gross-Schmetterlinge der Erde'' 13: Die Afrikanischen Tagfalter. Plate XIII 31

External links

Images of ''C. p. bebra''

Royal Museum for Central Africa The Royal Museum for Central Africa or RMCA ( nl, Koninklijk Museum voor Midden-Afrika or KMMA; french: Musée royal de l'Afrique centrale or MRAC; german: Königliches Museum für Zentralafrika or KMZA), also officially known as the AfricaMuse ...

(Albertine Rift Project)
Images of ''C. p. delvauxi''
(Albertine Rift Project)
Images of ''C. p. pleione ''
(Albertine Rift Project)
''C. p. pleione'' images
at

Consortium for the Barcode of Life The Consortium for the Barcode of Life (CBOL) was an international initiative dedicated to supporting the development of DNA barcoding as a global standard for species identification. CBOL's Secretariat Office is hosted by the National Museum of ...

''C. p. bebra'' images
at BOLD
''C. p. congoensis'' images
at BOLD
''C. p. delvauxi'' images
at BOLD Butterflies described in 1824 pleione {{Charaxinae-stub