butterfly
Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
in the family
Nymphalidae
The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to the superfamily Papilionoidea, they are usually medium-sized to large butterflies. Most species have a red ...
. It is found in north-eastern
Madagascar
Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
'', but has the tails on the hindwing much longer,8—9 mm. in length; the basal part of the upper surface is light red-brown and but little darker than the broad orange-brown median band; the cell of the forewing with two large black spots and the median band with large black spots in cellules 2—4; the hindwing above at the distal margin with orange-yellow transverse spots in 2—7; the hindwing beneath, with three black, white-bordered transverse spots at the costal margin in cellule 8;the ground-colour of the basal part bright red-brown. Madagascar; rare. Aurivillius, .O.. 1908-1924. In: Seitz, A. ''Die Großschmetterlinge der Erde'' Band 13: Abt. 2, ''Die exotischen Großschmetterlinge, Die afrikanischen Tagfalter'', 1925, 613 Seiten, 80 Tafeln (The Macrolepidoptera of the World 13).Alfred Kernen Verlag, Stuttgart.
Related species
Historical attempts to assemble a cluster of presumably related species into a "''Charaxes jasius'' Group" have not been wholly convincing. More recent taxonomic revision, corroborated by phylogenetic research, allow a more rational grouping congruent with cladistic relationships. Within a well-populated
clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
of 27 related species sharing a common ancestor approximately 16 mya during the
Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
, 26 are now considered together as The ''jasius'' Group. One of the two lineages within this clade forms a robust
monophyletic
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
group of seven species sharing a common ancestor approximately 2-3 mya, i.e. during the
Pliocene
The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58"Out of Africa again: A phylogenetic hypothesis of the genus ''Charaxes (Lepidoptera: Nymphalidae)'' based on five gene regions" Aduse-Poku, Vingerhoedt, Wahlberg. Molecular Phylogenetics and Evolution (2009) 53;463–478 and are considered as the ''jasius'' subgroup. The second lineage leads to 19 other species within the Jasius group, which are split in to three well-populated subgroups of closely related species.
The ''jasius'' Group (26 Species):
Clade 1: ''jasius'' subgroup (7 species)
Clade 2: contains the well-populated three additional subgroups (19 species) of the jasius Group: called the ''brutus'', ''pollux'', and ''eudoxus'' subgroups.
*the ''eudoxus'' subgroup (11 species):
*''
Charaxes eudoxus
__NOTOC__
''Charaxes eudoxus'', the Eudoxus charaxes, is a butterfly in the family Nymphalidae. It is found in Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, Nigeria, Cameroon, Equatorial Guinea, Gabon, the Republic of the Congo, Angola, the ...
Charaxes richelmanni
__NOTOC__
''Charaxes richelmanni'' is a butterfly in the family Nymphalidae. It is found in Cameroon, Gabon, the Republic of Congo, the Central African Republic, the Democratic Republic of Congo and Tanzania. The habitat consists of lowland eve ...
Charaxes andranodorus
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''Charaxes andranodorus'' is a butterfly in the family Nymphalidae. It is found on Madagascar.
Description
This rare species is one of the most magnificent forms of ''Charaxes''. Above it agrees very nearly with ''phraortes'', the bas ...
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*''
Charaxes andrefana
The rajah and pasha butterflies, also known as emperors in Africa and Australia, (genus ''Charaxes'') make up the huge type genus of the brush-footed butterfly subfamily Charaxinae, or leafwing butterflies. They belong to the tribe Charaxini, w ...
'' tat.rev.2025ref name="Turlin 2005"/>
Further exploration of the phylogenetic relationships amongst existing ''Charaxes'' taxa is required to improve clarity.
References
*
Victor Gurney Logan Van Someren
Victor Gurney Logan Van Someren (1886 in Melbourne – 24 July 1976) was a zoologist and entomologist.
Van Someren was born in Australia. He attended George Watson's College and studied zoology at University of Edinburgh. He was also a dentist ...
, 1974 Revisional notes on African ''Charaxes'' (Lepidoptera: Nymphalidae). Part IX. ''Bulletin of the British Museum of Natural History'' (Entomology) 29 (8):415-487
Consortium for the Barcode of Life
The Consortium for the Barcode of Life (CBOL) was an international initiative dedicated to supporting the development of DNA barcoding as a global standard for species identification. CBOL's Secretariat Office is hosted by the National Museum of ...
phraortes
Phraortes ( peo, 𐎳𐎼𐎺𐎼𐎫𐎡𐏁, translit=Fravartiš; grc, Φραόρτης, translit=Phraórtēs; died c. 653 BC), son of Deioces, was the second king of the Median Empire.
Like his father Deioces, Phraortes started wars agains ...