Charaxes Etesipe
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''Charaxes etesipe'', the savannah charaxes or scarce forest emperor, is a
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
of the family
Nymphalidae The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to the superfamily Papilionoidea, they are usually medium-sized to large butterflies. Most species have a red ...
.


Description

The
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan of ...
is for male. The females are larger, reaching . The uppersides of the forewings are blackish blue with a greenish tint. The outer margin are generally denticulate (tooth like), with a series of white discal spots. In the hindwings of both sexes there are short tails and a complete series of large pale blue patches. The undersides of the wings are variegated with cream and drab colours. A full description is given by Rothschild, W. And Jordan, K., 1900 ''Novitates Zoologicae'' Volume 7:455-45

page 428-429 (for terms see ''Novitates Zoologicae'' Volume 5:545-601

Seitz-The transverse band on the upperside of the hindwing in the male is blue and placed beyond the middle of the wing, in the female it is at least partly white and placed about at the middle. — ''etesipe'' Godt. male The transverse band of the forewing is narrow, entirely broken up into spots, only those in cellules la, lb and 2 are blue, the others white, and placed near the distal margin; white spots are also present in the basal part of cellules 2—6 and at the apex of the cell; the median band on the upperside of the hindwing is only 2–3 mm. in breadth and is also broken up into spots. Female- The transverse band on the upper surface is broad and white, the markings at the distal margin of the hindwing indistinct. Sierra Leone to Angola and Unyoro. — ''abyssinicus'' Rothsch. differs from the type-form in having the blue transverse band of the hindwing in the male continuous and about 5 mm. in breadth; the blue spot in cellules la and lb on the upperside of the forewing are also larger than in ''etesipe'' ; the female is unknown. Abyssinia. — ''tavetensis'' Rothsch. Male : he blue transverse band on the upperside of the hindwing is posteriorly continuous and very broad (7—8mm. in breadth in cellule 2), but anteriorly much narrowed and broken up into spots; the blue spots in cellules la and 1 b of the forewing are larger than in ''etesipe'', but smaller than in ''abyssinicus''. The female is very similar to that of typical ''etesipe'', but somewhat smaller and with more distinct markings at the distal margin of the hindwing above. East Africa from Nyassaland to Taveta in British East Africa.


Biology

Adults are on wing from August to October and from March to June. There are two generations per year. The larvae of subspecies ''tavetensis'' feed on ''
Afzelia quanzensis ''Afzelia quanzensis'' (pod mahogany, af, peulmahonie, ve, mutokota, zu, inkehli) is a protected tree in South Africa. This deciduous tree can grow up to 20m in height, and has smooth, grey bark, which can flake in irregular patches. It has glo ...
'', ''
Dalbergia nitidula ''Dalbergia'' is a large genus of small to medium-size trees, shrubs and lianas in the pea family, Fabaceae, subfamily Faboideae. It was recently assigned to the informal monophyletic ''Dalbergia'' clade (or tribe): the Dalbergieae. The g ...
'', '' Securidaca longipeduncularis'' and ''
Margaritaria discoidea ''Margaritaria discoidea'' is a tree in the family Phyllanthaceae, commonly known as the pheasant-berry, egossa red pear or bushveld peacock-berry. These trees are native to the warmer, higher rainfall areas of Africa. Taxonomy This species has ...
''. Other subspecies have been recorded on ''
Entada ''Entada'' is a genus of flowering plants in the family Fabaceae, in the mimosoid clade of the subfamily Caesalpinioideae. It consists of some 30 species of trees, shrubs and tropical lianas. About 21 species are known from Africa, six from A ...
'' and ''
Afzelia quanzensis ''Afzelia quanzensis'' (pod mahogany, af, peulmahonie, ve, mutokota, zu, inkehli) is a protected tree in South Africa. This deciduous tree can grow up to 20m in height, and has smooth, grey bark, which can flake in irregular patches. It has glo ...
''. Notes on the biology of ''etesipe'' are given by Kielland, J. (1990) and Larsen, T.B. (2005)


Distribution

This species can be found in tropical east and west Africa and in
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
in a mosaic of habitats but mainly in Afrotropical forest.


Taxonomy

Kielland regards to be closely related to ''
Charaxes penricei ''Charaxes penricei'', the scarce savanna charaxes, is a butterfly in the family Nymphalidae. It is found in the Republic of the Congo, Angola, the Democratic Republic of the Congo, Zambia, Tanzania, Malawi and Zimbabwe. Description The origina ...
'' and ''
Charaxes cacuthis ''Charaxes cacuthis'' is a butterfly in the family Nymphalidae. It is found on Madagascar. The habitat consists of lowland and coastal forests. Charaxes cacuthis up290209.jpg , recto Charaxes cacuthis down290209.jpg , verso Description A full ...
'' and considers ''tavetensis'' as a species distinct from ''etesipe'' and regards ''pemba'' and ''shaba'' as subspecies of ''tavetensis'' . ''Charaxes etesipe'' is a member of the
species group In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
''Charaxes etesipe'' The
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
members are: *''Charaxes etesipe'' nominate *''
Charaxes penricei ''Charaxes penricei'', the scarce savanna charaxes, is a butterfly in the family Nymphalidae. It is found in the Republic of the Congo, Angola, the Democratic Republic of the Congo, Zambia, Tanzania, Malawi and Zimbabwe. Description The origina ...
'' *''
Charaxes achaemenes ''Charaxes achaemenes'', the bushveld emperor or bush charaxes, is a butterfly of the family Nymphalidae found across Africa.Charaxes paradoxa ''Charaxes paradoxa'' is a butterfly in the family Nymphalidae. It is found on Grande Comore. The habitat consists of forests. The larvae feed on ''Phyllanthus'' species. Taxonomy ''Charaxes paradoxa'' is a member of the species group ''Charaxe ...
'' *''
Charaxes cacuthis ''Charaxes cacuthis'' is a butterfly in the family Nymphalidae. It is found on Madagascar. The habitat consists of lowland and coastal forests. Charaxes cacuthis up290209.jpg , recto Charaxes cacuthis down290209.jpg , verso Description A full ...
'' *''
Charaxes bwete The rajah and pasha butterflies, also known as emperors in Africa and Australia, (genus ''Charaxes'') make up the huge type genus of the brush-footed butterfly subfamily Charaxinae, or leafwing butterflies. They belong to the tribe Charaxini, w ...
'' *''
Charaxes cristalensis The rajah and pasha butterflies, also known as emperors in Africa and Australia, (genus ''Charaxes'') make up the huge type genus of the brush-footed butterfly subfamily Charaxinae, or leafwing butterflies. They belong to the tribe Charaxini, w ...
''


Subspecies

*''C. e. etesipe'' (
Sierra Leone Sierra Leone,)]. officially the Republic of Sierra Leone, is a country on the southwest coast of West Africa. It is bordered by Liberia to the southeast and Guinea surrounds the northern half of the nation. Covering a total area of , Sierra ...
,
Ivory Coast Ivory Coast, also known as Côte d'Ivoire, officially the Republic of Côte d'Ivoire, is a country on the southern coast of West Africa. Its capital is Yamoussoukro, in the centre of the country, while its largest city and economic centre is ...
,
Liberia Liberia (), officially the Republic of Liberia, is a country on the West African coast. It is bordered by Sierra Leone to Liberia–Sierra Leone border, its northwest, Guinea to its north, Ivory Coast to its east, and the Atlantic Ocean ...
,
Benin Benin ( , ; french: Bénin , ff, Benen), officially the Republic of Benin (french: République du Bénin), and formerly Dahomey, is a country in West Africa. It is bordered by Togo to the west, Nigeria to the east, Burkina Faso to the north ...
,
Togo Togo (), officially the Togolese Republic (french: République togolaise), is a country in West Africa. It is bordered by Ghana to the west, Benin to the east and Burkina Faso to the north. It extends south to the Gulf of Guinea, where its c ...
,
Ghana Ghana (; tw, Gaana, ee, Gana), officially the Republic of Ghana, is a country in West Africa. It abuts the Gulf of Guinea and the Atlantic Ocean to the south, sharing borders with Ivory Coast in the west, Burkina Faso in the north, and To ...
,
Nigeria Nigeria ( ), , ig, Naìjíríyà, yo, Nàìjíríà, pcm, Naijá , ff, Naajeeriya, kcg, Naijeriya officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf o ...
, southern
Chad Chad (; ar, تشاد , ; french: Tchad, ), officially the Republic of Chad, '; ) is a landlocked country at the crossroads of North and Central Africa. It is bordered by Libya to the north, Sudan to the east, the Central African Republic ...
,
Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...
, southern
Sudan Sudan ( or ; ar, السودان, as-Sūdān, officially the Republic of the Sudan ( ar, جمهورية السودان, link=no, Jumhūriyyat as-Sūdān), is a country in Northeast Africa. It shares borders with the Central African Republic t ...
, northern
Angola , national_anthem = " Angola Avante"() , image_map = , map_caption = , capital = Luanda , religion = , religion_year = 2020 , religion_ref = , coordina ...
,
Uganda }), is a landlocked country in East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territor ...
, northern
Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...
) *''C. e. abyssinicus'' Rothschild & Jordan, 1900 (
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
(west of Rift) *''C. e. gordoni'' van Someren, 1935 (
Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...
(north-eastern side of Mount Kenya)) *''C. e. hercules'' Turlin & Lequeux, 2002 (
Uganda }), is a landlocked country in East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territor ...
) *''C. e. patrizii'' Storace, 1949 (
Somalia Somalia, , Osmanya script: 𐒈𐒝𐒑𐒛𐒐𐒘𐒕𐒖; ar, الصومال, aṣ-Ṣūmāl officially the Federal Republic of SomaliaThe ''Federal Republic of Somalia'' is the country's name per Article 1 of thProvisional Constituti ...
) *''C. e. pemba'' van Someren, 1966 . (
Pemba Island Pemba Island ( ar, الجزيرة الخضراء ''al-Jazīra al-khadrā'', literally "The Green Island"; sw, Pemba kisiwa) is a Tanzanian island forming part of the Zanzibar Archipelago, lying within the Swahili Coast in the Indian Ocean. Geog ...
) *''C. e. shaba'' Berger, 1981 (
Zaire Zaire (, ), officially the Republic of Zaire (french: République du Zaïre, link=no, ), was a Congolese state from 1971 to 1997 in Central Africa that was previously and is now again known as the Democratic Republic of the Congo. Zaire was, ...
(Shaba)) *''C. e. tavetensis'' Rothschild, 1894Rothschild, W. 1894. Some new species of Lepidoptera. ''
Novitates Zoologicae ''Novitates Zoologicae: A Journal of Zoology in Connection With the Tring Museum'' was a British scientific journal devoted to systematic zoology. It was edited by Lionel Walter Rothschild and published between 1894 and 1948 by the Tring Museum ...
'' 1: 535-540.
(
Malawi Malawi (; or aláwi Tumbuka: ''Malaŵi''), officially the Republic of Malawi, is a landlocked country in Southeastern Africa that was formerly known as Nyasaland. It is bordered by Zambia to the west, Tanzania to the north and northeast ...
,
Mozambique Mozambique (), officially the Republic of Mozambique ( pt, Moçambique or , ; ny, Mozambiki; sw, Msumbiji; ts, Muzambhiki), is a country located in southeastern Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi ...
, coast of Kenya,
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...
,
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
) perhaps a species


Subspecies gallery

File:Nymphalidae - Charaxes etesipe.JPG, ''C. e. abyssinicus'' from Ethiopia File:Charaxes etesipe recto.jpg, ''C. e. etesipe'' from Cameroon - dorsal view File:Charaxes etesipe verso.jpg, ''C. e. etesipe'' from Cameroon - ventral view


References

*
Victor Gurney Logan Van Someren Victor Gurney Logan Van Someren (1886 in Melbourne – 24 July 1976) was a zoologist and entomologist. Van Someren was born in Australia. He attended George Watson's College and studied zoology at University of Edinburgh. He was also a dentist ...
(1966). Revisional notes on African ''Charaxes'' (Lepidoptera: Nymphalidae). Part III. ''Bulletin of the British Museum'' (Natural History) (Entomology) 45–10


External links


Pteron World

"''Charaxes etesipe'' (Godart, 1824)"
''Insecta.pro''.
''Charaxes etesipe'' images
subspecies and forma at
Consortium for the Barcode of Life The Consortium for the Barcode of Life (CBOL) was an international initiative dedicated to supporting the development of DNA barcoding as a global standard for species identification. CBOL's Secretariat Office is hosted by the National Museum of ...
.
''Charaxes tavetensis'' images
at BOLD
''Charaxes tavetensis tavetensis'' images
at BOLD (verso)
''Charaxes tavetensis pemba'' images
at BOLD
Images of ''C. etesipe etesipe''
Royal Museum for Central Africa The Royal Museum for Central Africa or RMCA ( nl, Koninklijk Museum voor Midden-Afrika or KMMA; french: Musée royal de l'Afrique centrale or MRAC; german: Königliches Museum für Zentralafrika or KMZA), also officially known as the AfricaMuse ...
(Albertine Rift Project)
Images of ''C. etesipe gordoni''
(Albertine Rift Project)
Images of ''C. etesipe shaba''
(Albertine Rift Project)
Images of ''C. etesipe tavetensis''
(Albertine Rift Project) {{Taxonbar , from=Q2957634 Butterflies described in 1824 etesipe