''Chaos'' is a
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
single-celled amoeboid
An amoeba (; less commonly spelled ameba or amœba; plural ''am(o)ebas'' or ''am(o)ebae'' ), often called an amoeboid, is a type of cell or unicellular organism with the ability to alter its shape, primarily by extending and retracting pseudopo ...
organisms
In biology, an organism () is any living system that functions as an individual entity. All organisms are composed of cells (cell theory). Organisms are classified by taxonomy into groups such as multicellular animals, plants, and fungi; ...
in the family
Amoebidae
The Amoebidae are a family of Amoebozoa, including naked amoebae that produce multiple pseudopodia of indeterminate length. These are roughly cylindrical with granular endoplasm and no subpseudopodia, as found in other members of the class Tubul ...
. The largest and best-known species, the so-called "giant amoeba" ''Chaos carolinensis'', can reach lengths of 5 mm, although most specimens fall between 1 and 3 mm.
Members of the genus closely resemble those of genus ''
Amoeba
An amoeba (; less commonly spelled ameba or amœba; plural ''am(o)ebas'' or ''am(o)ebae'' ), often called an amoeboid, is a type of Cell (biology), cell or unicellular organism with the ability to alter its shape, primarily by extending and ret ...
'' and share the same general morphology, producing numerous cylindrical
pseudopod
A pseudopod or pseudopodium (plural: pseudopods or pseudopodia) is a temporary arm-like projection of a eukaryotic cell membrane that is emerged in the direction of movement. Filled with cytoplasm, pseudopodia primarily consist of actin filament ...
s, each of which is rounded at the tip. However, while ''Amoeba'' have a single
nucleus
Nucleus ( : nuclei) is a Latin word for the seed inside a fruit. It most often refers to:
*Atomic nucleus, the very dense central region of an atom
* Cell nucleus, a central organelle of a eukaryotic cell, containing most of the cell's DNA
Nucl ...
, ''Chaos'' can have as many as a thousand. Because of this attribute, ''C. carolinensis'' was once placed in the genus ''
Pelomyxa
''Pelomyxa'' is a genus of giant flagellar amoebae, usually 500-800 μm but occasionally up to 5 mm in length, found in anaerobic or microaerobic bottom sediments of stagnant freshwater ponds or slow-moving streams.Chistyakova, L. V., and A. ...
'' alongside the giant
multinucleate
Multinucleate cells (also known as multinucleated or polynuclear cells) are eukaryotic cells that have more than one nucleus per cell, i.e., multiple nuclei share one common cytoplasm. Mitosis in multinucleate cells can occur either in a coordinat ...
amoeba ''Pelomyxa palustris.'' Recently, molecular phylogenetic studies of this species have confirmed the view of some earlier researchers that it is more closely related to ''Amoeba'' than to ''Pelomyxa''. The species is now placed in the independent genus ''Chaos'', a sister group to ''Amoeba''.
Dietary habits
''Chaos'' species are versatile
heterotrophs
A heterotroph (; ) is an organism that cannot produce its own food, instead taking nutrition from other sources of organic carbon, mainly plant or animal matter. In the food chain, heterotrophs are primary, secondary and tertiary consumers, but ...
, able to feed on bacteria, algae, other protists, and even small
multicellular
A multicellular organism is an organism that consists of more than one cell, in contrast to unicellular organism.
All species of animals, land plants and most fungi are multicellular, as are many algae, whereas a few organisms are partially uni- ...
invertebrate
Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s.
Like all
Amoebozoa
Amoebozoa is a major taxonomic group containing about 2,400 described species of amoeboid protists, often possessing blunt, fingerlike, lobose pseudopods and tubular mitochondrial cristae. In traditional and currently no longer supported classi ...
, they take in food by
phagocytosis
Phagocytosis () is the process by which a cell uses its plasma membrane to engulf a large particle (≥ 0.5 μm), giving rise to an internal compartment called the phagosome. It is one type of endocytosis. A cell that performs phagocytosis is ...
, encircling food particles with its pseudopodia, then enclosing them within a food ball, or
vacuole
A vacuole () is a membrane-bound organelle which is present in plant and fungal cells and some protist, animal, and bacterial cells. Vacuoles are essentially enclosed compartments which are filled with water containing inorganic and organic mo ...
, where they are broken down by enzymes. The cell does not have a mouth or
cytostome
A cytostome (from ''cyto-'', cell and ''stome-'', mouth) or cell mouth is a part of a cell specialized for phagocytosis, usually in the form of a microtubule-supported funnel or groove. Food is directed into the cytostome, and sealed into vacuol ...
, nor is there any fixed site on the cell membrane at which phagocytosis normally occurs.
Movement
The cell's membrane, or
plasmalemma
The cell membrane (also known as the plasma membrane (PM) or cytoplasmic membrane, and historically referred to as the plasmalemma) is a biological membrane that separates and protects the interior of all cells from the outside environment (th ...
, is extremely flexible, allowing the organism to change shape from one moment to the next. The
cytoplasm
In cell biology, the cytoplasm is all of the material within a eukaryotic cell, enclosed by the cell membrane, except for the cell nucleus. The material inside the nucleus and contained within the nuclear membrane is termed the nucleoplasm. The ...
within the membrane is conventionally described as having two parts: the internal fluid, or
endoplasm
Endoplasm generally refers to the inner (often granulated), dense part of a cell's cytoplasm. This is opposed to the ectoplasm which is the outer (non-granulated) layer of the cytoplasm, which is typically watery and immediately adjacent to the ...
, which contains loose granules and food vacuoles, as well as organelles such as nuclei and
mitochondria
A mitochondrion (; ) is an organelle found in the Cell (biology), cells of most Eukaryotes, such as animals, plants and Fungus, fungi. Mitochondria have a double lipid bilayer, membrane structure and use aerobic respiration to generate adenosi ...
; and a more viscous
ectoplasm around the perimeter of the cell, which is relatively clear and contains no conspicuous granules. Like other lobose amoebae, ''Chaos'' move by extending
pseudopod
A pseudopod or pseudopodium (plural: pseudopods or pseudopodia) is a temporary arm-like projection of a eukaryotic cell membrane that is emerged in the direction of movement. Filled with cytoplasm, pseudopodia primarily consist of actin filament ...
ia. As a new pseudopod is extended, a variable zone of ectoplasm forms at the leading edge and a fountaining stream of endoplasm circulates within. The effort of describing these motions, and explaining how they result in the cell's forward movement, has generated a large body of scientific literature.
Early history and naming controversy
![Amoebas from Ehrenberg](https://upload.wikimedia.org/wikipedia/commons/7/7b/Amoebas_from_Ehrenberg.jpg)
The genus ''Chaos'' has had a long and often confusing history. In 1755,
Rösel von Rosenhof saw and depicted an amoeboid he named "''der kleine Proteus''" ("the little Proteus"). Three years later, Linnaeus gave Rösel's creature the name ''Volvox chaos''. However, because the name ''
Volvox
''Volvox'' is a polyphyletic genus of chlorophyte green algae in the family Volvocaceae. It forms spherical colonies of up to 50,000 cells. They live in a variety of freshwater habitats, and were first reported by Antonie van Leeuwenhoek in 1700 ...
'' had already been applied to a genus of flagellate algae, he later changed it to ''Chaos chaos''. In subsequent decades, as new names and species proliferated, accounts of ''Chaos'', under a variety of synonyms, became so thoroughly entangled with descriptions of similar organisms, that it is virtually impossible to differentiate one historic amoeboid from another. In 1879, Joseph Leidy suggested collapsing all the "common" large, freshwater amoebae into one species, which he proposed to call ''
Amoeba proteus
''Amoeba proteus'' is a large species of amoeba closely related to another genus of giant amoebae, ''Chaos''. As such, the species is sometimes given the alternative scientific name ''Chaos diffluens''.
This protozoan uses extensions called pseu ...
''. A dozen species, including several that had been identified as belonging to ''Chaos'', were to be regarded as synonyms of ''Amoeba proteus''. However, in the description he gives of this organism, it is clearly defined as a