The vagabond butterflyfish (''Chaetodon vagabundus''), also known as the crisscross butterflyfish, is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

of marine ray-finned fish, a

butterflyfish The butterflyfish are a group of conspicuous tropical marine fish of the family Chaetodontidae; the bannerfish and coralfish are also included in this group. The approximately 129 species in 12 genera are found mostly on the reefs of the Atlant ...

belonging to the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Chaetodontidae. It is found in the

Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...

region.

Description

The vagabond butterflyfish has a whitish body which is marked with two series of thin dark diagonal lines perpendicular to each other, forming a

chevron Chevron (often relating to V-shaped patterns) may refer to: Science and technology * Chevron (aerospace), sawtooth patterns on some jet engines * Chevron (anatomy), a bone * ''Eulithis testata'', a moth * Chevron (geology), a fold in rock lay ...

pattern. There is also a wide black vertical band running through the eye and a second band running through the caudal peduncle and a third on the centre of the

caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

. There are very thin orange horizontal lines over the forehead. The juveniles have a black dot on the soft-rayed part of their

dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...

, near the posterior end. The dorsal fin contains 13 spines and 23-25 soft rays while the anal fin contains 3 spines and 19-22 soft rays. This species grows to a maximum total length of although a more typical length would be .

Distribution

The vagabond butterflyfish is found in the Indian and Pacific Oceans. It occurs from the eastern coast of Africa, where it is found from

Socotra Socotra or Soqotra (; ar, سُقُطْرَىٰ ; so, Suqadara) is an island of the Republic of Yemen in the Indian Ocean, under the ''de facto'' control of the UAE-backed Southern Transitional Council, a secessionist participant in Yemen’s ...

KwaZulu-Natal KwaZulu-Natal (, also referred to as KZN and known as "the garden province") is a province of South Africa that was created in 1994 when the Zulu bantustan of KwaZulu ("Place of the Zulu" in Zulu) and Natal Province were merged. It is locate ...

in South Africa; across the Indian Ocean east as far as the Line and Gambier Islands in Polynesia, north to southern Japan and south to

New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...

and Rapa Iti.

Habitat and biology

The vagabond butterflyfish is found in reef areas including coastal reef flats within the reef, lagoons and more exposed outer reef slopes. It can tolerate a variety of environmental conditions, such as turbid waters and freshwater plumes in the vicinity of the mouths of streams. They are omnivorous, known to feed on

algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...

coral polyp Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and sec ...

crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...

s and

worm Worms are many different distantly related bilateral animals that typically have a long cylindrical tube-like body, no limbs, and no eyes (though not always). Worms vary in size from microscopic to over in length for marine polychaete wor ...

s. These

oviparous Oviparous animals are animals that lay their eggs, with little or no other embryonic development within the mother. This is the reproductive method of most fish, amphibians, most reptiles, and all pterosaurs, dinosaurs (including birds), and ...

monogamous Monogamy ( ) is a form of Dyad (sociology), dyadic Intimate relationship, relationship in which an individual has only one Significant other, partner during their lifetime. Alternately, only one partner at any one time (Monogamy#Serial monogamy, ...

fish form stable pairs with both pair members jointly defending a feeding

territory A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...

against other pairs. However they often accompany other species without being aggressive. By the standards of their genus, they are easily maintained in tanks.

Systematics

The vagabond butterflyfish was first formally described in 1758 by Carl Linnaeus in the 10th edition of his ''

Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...

''. It belongs to the large

subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...

''Rabdophorus'' which might warrant recognition as a distinct

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

. In this group, it almost certainly is a rather close relative of the

threadfin butterflyfish The threadfin butterflyfish (''Chaetodon auriga'') is a species of marine ray-finned fish belonging to the family (biology), family Chaetodontidae. It has a wide Indo-Pacific distribution. Distribution ''Chaetodon auriga'' is found in the Indo-P ...

(''C. auriga'') and the

Indian vagabond butterflyfish The Indian vagabond butterflyfish (''Chaetodon decussatus''), also known as the blackened butterflyfish or black-finned vagabond, is a species of marine ray-finned fish, a butterflyfish belonging to the family Chaetodontidae. It is found in the ...

(''C. decussatus''). The latter might be closer to the threadfin butterflyfish; as ''C. vagabundus'' has yielded abnormal mtDNA

12S rRNA Mitochondrially encoded 12S ribosomal RNA (often abbreviated as 12S or 12S rRNA), also known as Mitochondrial-derived peptide MOTS-c or Mitochondrial open reading frame of the 12S rRNA-c is the SSU rRNA of the mitochondrial ribosome. In humans, ...

sequence In mathematics, a sequence is an enumerated collection of objects in which repetitions are allowed and order matters. Like a set, it contains members (also called ''elements'', or ''terms''). The number of elements (possibly infinite) is calle ...

data this is hard to say however. The ''C. auriga'' species group shares the characteristic pattern of two areas of ascending and descending oblique lines; species differ conspicuously in hindpart coloration.

References

External links

* {{Taxonbar, from=Q389163 Chaetodon Fish of the Red Sea Fish described in 1758 Taxa named by Carl Linnaeus