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Cetotheriidae is a
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
of
baleen whale Baleen whales (systematic name Mysticeti), also known as whalebone whales, are a parvorder of carnivorous marine mammals of the infraorder Cetacea (whales, dolphins and porpoises) which use keratinaceous baleen plates (or "whalebone") in their ...
s (parvorder
Mysticeti Baleen whales (systematic name Mysticeti), also known as whalebone whales, are a parvorder of carnivorous marine mammals of the infraorder Cetacea (whales, dolphins and porpoises) which use keratinaceous baleen plates (or "whalebone") in their ...
). The family is known to have existed from the
Late Oligocene The Chattian is, in the geologic timescale, the younger of two ages or upper of two stages of the Oligocene Epoch/Series. It spans the time between . The Chattian is preceded by the Rupelian and is followed by the Aquitanian (the lowest stage ...
to the
Early Pleistocene The Early Pleistocene is an unofficial sub-epoch in the international geologic timescale in chronostratigraphy, being the earliest division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently estimated to span the time ...
before going
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
. Although some
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
studies conducted by recovered the living
pygmy right whale The pygmy right whale (''Caperea marginata'') is a species of baleen whale. It may be a member of the cetotheres, a family of baleen whales which until 2012 were thought to be extinct; ''C. marginata'' has otherwise been considered the monotyp ...
as a member of Cetotheriidae, making the pygmy right whale the only living cetotheriid, other authors either dispute this placement or recover Neobalaenidae as a sister group to Cetotheriidae.


Taxonomy

After its description by Brant in 1872, Cetotheriidae was used as a
wastebasket taxon Wastebasket taxon (also called a wastebin taxon, dustbin taxon or catch-all taxon) is a term used by some taxonomists to refer to a taxon that has the sole purpose of classifying organisms that do not fit anywhere else. They are typically defined ...
for baleen whales which were not assignable to
extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...
whale families. Comparing the cranial and mandibular morphology of 23 taxa (including late archaeocetes and both fossil and extant mysticetes), found Cetotheriidae in this traditional sense to be polyphyletic. Based on ten cranial characters, they also concluded that of the twelve included fossil baleen-bearing mysticetes, six formed a monophyletic group, Cetotheriidae ''
sensu stricto ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular co ...
''. Several phylogenetic studies since Bouetel & Muizon 2006 support the monophyly of a small group of core Cetotheriidae ''sensu stricto'', archaic mysticetes with a cranium that have "a long ascending process of the maxilla with anteriorly diverging lateral border that interdigitates with the frontal" and some other characters. This group is limited to ''Cetotherium rathkii'', ''Metopocetus durinasus'', ''Mixocetus elysius'', ''Herpetocetus scaldiensis'', ''H. transatlanticus'', ''H. bramblei'', ''Nannocetus eremus'', and ''Piscobalaena nana''. The remaining genera placed in the family are considered Cetotheriidae ''
sensu lato ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular co ...
'' and are often referred to as the 'cetotheres'. considered the primitive 'cetothere' ''
Joumocetus ''Joumocetus'' is a genus of extinct baleen whale in the family Cetotheriidae containing the single species ''Joumocetus shimizui''. The species is known only from a partial skeleton found in Miocene age sediments of Japan. History and classifi ...
'' the most basal named taxon of their new superfamily Thalassotherii (Cetotheriidae ''s.l.'', Cetotheriidae ''s.s.'',
Eschrichtiidae Eschrichtiidae or the gray whales is a family of baleen whale (Parvorder Mysticeti) with a single extant species, the gray whale (''Eschrichtius robustus''), as well as three described fossil genera: '' Archaeschrichtius'' and ''Eschrichtioide ...
(gray whales) and Balaenopteridae (rorquals)) and suggested that the term "Cetotheriidae ''s.l.''" should be renamed "basal" or "stem thalassotherians". found that the pygmy right whale formed a well-supported clade with Eschrichtiidae and Balaenopteridae based on molecular data, and that, within 'cetotheres', it was most closely related to the herpetocetines (''
Herpetocetus ''Herpetocetus'' is a genus of cetotheriid mysticete in the subfamily Herpetocetinae. Taxonomy There are four recognized species of ''Herpetocetus'': ''H. scaldiensis'', ''H. transatlanticus'',Whitmore, F.C., and L.G. Barnes. 2008. The Herpet ...
'' and ''
Nannocetus ''Nannocetus'' is an extinct baleen whale belonging to the family Cetotheriidae. Description ''Nannocetus'' is a diminutive mysticete measuring 13 feet (4 meters) long. It is characterized by the ventral orientation (in posterior view) of the p ...
'') Bisconti et al. 2013, however, found, based on morphological data, it to be more closely related to
Balaenidae Balaenidae () is a family of whales of the parvorder Mysticeti that contains two living genera: the right whales (genus ''Eubalaena''), and in a separate genus, the closely related bowhead whale (genus ''Balaena''). Evolutionary history Bale ...
(the bowhead and right whales), but added that additional specimens are expected to resolve these conflicting results within a few years. Classification of Cetotheriidae according to El Adli et al. (2014) and the Fossilworks online database: * †'' Cephalotropis'' Cope 1896 * †''
Joumocetus ''Joumocetus'' is a genus of extinct baleen whale in the family Cetotheriidae containing the single species ''Joumocetus shimizui''. The species is known only from a partial skeleton found in Miocene age sediments of Japan. History and classifi ...
'' * Subfamily: † Cetotheriinae Whitmore and Barnes, 2008 ** †''
Brandtocetus ''Brandtocetus'' is a genus of cetotheriid mysticete in the subfamily Cetotheriinae. The type and only species is ''Brandtocetus chongulek'' from the late Miocene (Tortonian) of the Kerch Peninsula in Crimea. ''Brandtocetus chongulek'' was a wh ...
'' Gol'din and Startsev 2014 ** †''
Cetotherium ''Cetotherium'' ("whale beast") is an extinct genus of baleen whales from the family Cetotheriidae. Taxonomy The family Cetotheriidae and the genus ''Cetotherium'' (sensu lato) have been used as wastebaskets for all kinds of baleen whales, mo ...
'' Brandt 1843 ** †''
Ciuciulea Ciuciulea is a village in Glodeni District, Moldova.Eucetotherium ''Eucetotherium'' is a genus of cetotheriid mysticete from Miocene (Tortonian The Tortonian is in the geologic time scale an age or stage of the late Miocene that spans the time between 11.608 ± 0.005 Ma and 7.246 ± 0.005 Ma (million year ...
'' Brandt 1873 ** †''
Kurdalagonus ''Kurdalagonus'' is a genus of cetotheriid mysticete in the subfamily Cetotheriinae from the Miocene of the Russian Caucasus. ''"Kurdalagonus" adygeicus'' has been reassigned to ''Mithridatocetus'', while the referral of ''"Cetotherium ''Ceto ...
'' ** †''
Mithridatocetus ''Mithridatocetus'' is a genus of cetotheriid mysticete in the subfamily Cetotheriinae. Known specimens have been found in marine deposits in Crimea, Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is ...
'' Gol'din and Startsev 2017 ** †''
Vampalus ''Vampalus'' is a genus of cetotheriid mysticete from Miocene (Tortonian The Tortonian is in the geologic time scale an age or stage of the late Miocene that spans the time between 11.608 ± 0.005 Ma and 7.246 ± 0.005 Ma (million years ag ...
'' ** †''
Zygiocetus ''Zygiocetus'' is an extinct genus of cetotheriid mysticete in the subfamily Cetotheriinae. The type and only species is ''Zygiocetus nartorum'', known from the Late Miocene ( Messinian) of Adygea in the Russian Caucasus. ''Zygiocetus nartorum' ...
'' Tarasenko 2014 * Subfamily Herpetocetinae Steeman, 2007 ** †''
Herentalia ''Herentalia'' is a genus of cetotheriid Cetotheriidae is a family of baleen whales (parvorder Mysticeti). The family is known to have existed from the Late Oligocene to the Early Pleistocene before going extinct. Although some phylogenetic ...
'' Bisconti 2015 ** †''
Herpetocetus ''Herpetocetus'' is a genus of cetotheriid mysticete in the subfamily Herpetocetinae. Taxonomy There are four recognized species of ''Herpetocetus'': ''H. scaldiensis'', ''H. transatlanticus'',Whitmore, F.C., and L.G. Barnes. 2008. The Herpet ...
'' Van Beneden 1872 ** †'' Metopocetus'' Cope 1896 ** †''
Nannocetus ''Nannocetus'' is an extinct baleen whale belonging to the family Cetotheriidae. Description ''Nannocetus'' is a diminutive mysticete measuring 13 feet (4 meters) long. It is characterized by the ventral orientation (in posterior view) of the p ...
'' Kellogg 1929 ** †'' Piscobalaena'' Pilleri and Siber 1989 ** †''
Tranatocetus ''Tranatocetus'' is an extinct genus of mysticete from the late Miocene (Tortonian) of Jutland, Denmark. The type and only species is ''Tranatocetus argillarius''. Description ''Tranatocetus'' is similar to ''"Aulocetus" latus'', ''"Cetotheriu ...
''? Gol'din and Steeman 2015 ** †''
Miocaperea ''Miocaperea'' is an extinct genus of pygmy right whale from the Miocene, Late Miocene Pisco Formation of Peru.Caperea'' (Gray, 1846) ''Incertae sedis:'' * †'' Hibacetus'' Otsuka and Ota 2008


References


Notes


Sources

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External links

* * {{Authority control Baleen whales Mammal families Extant Chattian first appearances