Ceratophora Tennentii
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''Ceratophora tennentii'', commonly known as the rhinoceros agama, horn-nosed lizard, and Tennent's leaf-nosed lizard, is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Agamidae Agamidae is a family (biology), family of over 300 species of iguanian lizards indigenous to Africa, Asia, Australia, and a few in Southern Europe. Many species are commonly called dragons or dragon lizards. Overview phylogenetics, Phylogenetic ...
. The species is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
.


Etymology

The generic name, ''Ceratophora'', means horn bearer. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
, ''tennentii'', is in honour of Irish politician
James Emerson Tennent Sir James Emerson Tennent, 1st Baronet, FRS (born James Emerson; 7 April 1804 – 6 March 1869) was a British politician and traveller born in Ireland. He was elected a Fellow of the Royal Society on 5 June 1862. Life The third son of William ...
, who was Colonial Secretary of Ceylon (1845–1850).Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). ''The Eponym Dictionary of Reptiles''. Baltimore: Johns Hopkins University Press. xiii + 296 pp. . (''Ceratophora tennentii'', p. 263).


Description

''C. tennentii'' has a leaf-like appendage on the end of its nose. The species can reach lengths of over including the tail. Males usually have more green in their coloration than females have, although they can change color to a reddish brown. Females usually have a shorter appendage on the nose. ''C. tennentii'' is not very agile and relies more on coloration than speed to avoid predators. The head is oval, and longer than wide. The rostral appendage is fleshy, laterally compressed, leaf-like with a bluntly conical scale at the tip. The lamellae under fourth toe number 23-30. The dorsum is reddish brown to olive green. The larger flank scales are more green. The gular region and sides of the neck have dark markings. The tail has 20 dark brown cross-bands. The venter is cream-coloured.


Behaviour

''C. tennentii'' is diurnal, and therefore is usually only active in the day. Very little is actually known about the behaviour of this unusual lizard.


Habitat

''C. tennentii'' is found in the wet tropical montane
cloud forests A cloud forest, also called a water forest, primas forest, or tropical montane cloud forest (TMCF), is a generally tropical or subtropical, evergreen, montane, moist forest characterized by a persistent, frequent or seasonal low-level cloud ...
of the
Knuckles mountains The Knuckles Mountain Range lies in central Sri Lanka, in the Districts of Matale and Kandy. The range takes its name from a series of recumbent folds and peaks in the west of the massif which resemble the knuckles of clenched fist when viewed ...
in
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
at elevations of . It has also been recorded in several other forest habitats.


Diet

''C. tennentii'' is reported to feed on insects and other small arthropods.


Reproduction

''C. tennentii'' lays eggs and is a sexually reproducing animal.


Threats

Threats to ''C. tennentii'' include
deforestation Deforestation or forest clearance is the removal of a forest or stand of trees from land that is then converted to non-forest use. Deforestation can involve conversion of forest land to farms, ranches, or urban use. The most concentrated d ...
,
pesticides Pesticides are substances that are meant to control pests. This includes herbicide, insecticide, nematicide, molluscicide, piscicide, avicide, rodenticide, bactericide, insect repellent, animal repellent, microbicide, fungicide, and lampric ...
,
climate change In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to E ...
,
forest fire A wildfire, forest fire, bushfire, wildland fire or rural fire is an unplanned, uncontrolled and unpredictable fire in an area of Combustibility and flammability, combustible vegetation. Depending on the type of vegetation present, a wildfire ...
s, and
bioaccumulation Bioaccumulation is the gradual accumulation of substances, such as pesticides or other chemicals, in an organism. Bioaccumulation occurs when an organism absorbs a substance at a rate faster than that at which the substance is lost or eliminated ...
. Much of its habitat has been cleared for
illegal logging Illegal logging is the harvest, transportation, purchase or sale of timber in violation of laws. The harvesting procedure itself may be illegal, including using corrupt means to gain access to forests; extraction without permission, or from a pro ...
and the
cardamom Cardamom (), sometimes cardamon or cardamum, is a spice made from the seeds of several plants in the genera ''Elettaria'' and ''Amomum'' in the family Zingiberaceae. Both genera are native to the Indian subcontinent and Indonesia. They are rec ...
,
coffee Coffee is a drink prepared from roasted coffee beans. Darkly colored, bitter, and slightly acidic, coffee has a stimulant, stimulating effect on humans, primarily due to its caffeine content. It is the most popular hot drink in the world. S ...
,
tea Tea is an aromatic beverage prepared by pouring hot or boiling water over cured or fresh leaves of '' Camellia sinensis'', an evergreen shrub native to East Asia which probably originated in the borderlands of southwestern China and northe ...
, and
rubber Rubber, also called India rubber, latex, Amazonian rubber, ''caucho'', or ''caoutchouc'', as initially produced, consists of polymers of the organic compound isoprene, with minor impurities of other organic compounds. Thailand, Malaysia, and ...
plantations over the past two centuries.


Captivity

As of 2006, the leaf-nosed lizard is protected because of its endangered status, making its trade illegal.


Conservation efforts

In 2000, areas above above sea level were protected and labeled as conservation forest. Cardamom cultivation had to be abandoned in this area. However, rather than natural regeneration, the cardamom range was taken over by invasive weeds such as mistflower ('' Eupatorium riparium'') and lantana (''
Lantana camara ''Lantana camara'' (common lantana) is a species of flowering plant within the verbena family (Verbenaceae), native to the American tropics. It is a very adaptable species, which can inhabit a wide variety of ecosystems; once it has been introduc ...
''). ''C. tennentii'' was placed on the endangered list by the IUCN in 2006.


References


External links

*''Reptiles Magazine''

*Discovery Communications, LLC

Downloaded on 9 October 2012.


Further reading

* George Albert Boulenger, Boulenger GA (1887). ''Catalogue of the Lizards in the British Museum (Natural History). Second Edition. Volume I. ... Agamidæ.'' London: Trustees of the British Museum (Natural History). (Taylor and Francis, Printers). xii + 436 pp. + Plates I-XXXII. (''Ceratophora tennentii'', p. 278). *Boulenger GA (1890). ''The Fauna of British India, Including Ceylon and Burma. Reptilia and Batrachia''. London: Secretary of State for India in Council. (Taylor and Francis, printers). xviii + 541 pp. (''Ceratophora tennentii'', p. 120). * Günther A (1861). "''Ceratophora Tennentii'', new species". p. 281 + unnumbered plate. ''In'': Tennent JE (1861). ''Sketches of the Natural History of Ceylon with Narratives and Anecdotes Illustrative of the Habits and Instincts of the Mammalia, Birds, Reptiles, Fishes, Insects, &c. Including a Monograph of the Elephant and a Description of the Modes of Capturing and Training It''. London: Longman, Green, Longman & Roberts. xxiii + 500 pp. * Günther ACLG (1864). ''The Reptiles of British India''. London: The Ray Society. (Taylor and Francis, printers). xxvii + 452 pp. + Plates I-XXVI. (''Ceratophora tennentii'', pp. 130-131). * Smith MA (1935). ''The Fauna of British India, Including Ceylon and Burma. Reptilia and Amphibia. Vol. II.—Sauria.'' London: Secretary of State for India in Council. (Taylor and Francis, printers). xiii + 440 pp. + Plate I + 2 maps. ("''Ceratophora tennenti'' ic, pp. 153–154). {{Taxonbar, from=Q783468 Reptiles of Sri Lanka Ceratophora Lizards of Asia Endemic fauna of Sri Lanka Reptiles described in 1861 Taxa named by Albert Günther Taxonomy articles created by Polbot