''Ceratocapnos claviculata'', the climbing corydalis, is a weak scrambling plant in the family

Papaveraceae The Papaveraceae are an economically important family of about 42 genera and approximately 775 known species of flowering plants in the order Ranunculales, informally known as the poppy family. The family is cosmopolitan, occurring in temperat ...

. It is endemic to Europe, growing mostly near the Atlantic fringe.Botanical Society of Britain and Ireland

Description

This delicate looking plant is a hairless

annual Annual may refer to: *Annual publication, periodical publications appearing regularly once per year ** Yearbook ** Literary annual *Annual plant *Annual report *Annual giving *Annual, Morocco, a settlement in northeastern Morocco *Annuals (band), ...

(or occasionally

perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...

) up to a metre tall with weak, often pinkish, clambering stems. The leaves are pale to medium green, doubly compound, the leaflets being well-stalked and divided into three to five sub-leaflets, and ending in a branching tendril. It blooms between May and September in the UK. The flowers are small, about long, pale creamy-yellow, in short axilliary spikes. Each flower is elongated and tubular with a lip and spur and stamens in two bundles. The seed pods are short, usually narrowing between the two seeds.

Taxonomy

It was first published as ''Ceratocapnos claviculata'' by

Magnus Lidén Magnus Lidén (born 1951) is a Swedish systematic botanist. Lidén received a PhD in systematic botany from Gothenburg University in 1986, where he stayed until 1997. He was the director of the Uppsala Botanic Gardens from 1998 – 2003. Since 20 ...

in ' Anales Jard. Bot. Madrid' vol.41 on page 221 in 1984, based on an earlier description by

Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...

under the name ''Fumaria claviculata''. The Latin

specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...

''claviculata'' refers to having tendrils or being tendrilled.

Distribution

This species is known from several countries in western Europe,

Belgium Belgium, ; french: Belgique ; german: Belgien officially the Kingdom of Belgium, is a country in Northwestern Europe. The country is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeast, France to th ...

Denmark ) , song = ( en, "King Christian stood by the lofty mast") , song_type = National and royal anthem , image_map = EU-Denmark.svg , map_caption = , subdivision_type = Sovereign state , subdivision_name = Danish Realm, Kingdom of Denmark ...

France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...

Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...

Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...

Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...

Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...

Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...

and

Norway Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and t ...

. The largest proportion of the global population is found in the United Kingdom. It grows in most counties in Britain especially the more western ones, but is absent from

Orkney Orkney (; sco, Orkney; on, Orkneyjar; nrn, Orknøjar), also known as the Orkney Islands, is an archipelago in the Northern Isles of Scotland, situated off the north coast of the island of Great Britain. Orkney is 10 miles (16 km) north ...

Shetland Shetland, also called the Shetland Islands and formerly Zetland, is a subarctic archipelago in Scotland lying between Orkney, the Faroe Islands and Norway. It is the northernmost region of the United Kingdom. The islands lie about to the no ...

and the

Outer Hebrides The Outer Hebrides () or Western Isles ( gd, Na h-Eileanan Siar or or ("islands of the strangers"); sco, Waster Isles), sometimes known as the Long Isle/Long Island ( gd, An t-Eilean Fada, links=no), is an island chain off the west coast ...

and rare in

. UK conservation status is least concern as of 2005.

Ecology

Climbing corydalis tends to grow on the edges of woodlands and previously wooded sites. It prefers acid soils, sandy or peaty, and usually in sheltered and half shaded positions. It is sometimes abundant in disturbed parts of recently cleared plantations or woods, clambering over wood debris. It grows well in impoverished soil under

bracken Bracken (''Pteridium'') is a genus of large, coarse ferns in the family Dennstaedtiaceae. Ferns (Pteridophyta) are vascular plants that have alternating generations, large plants that produce spores and small plants that produce sex cells (eggs ...

, perhaps because it flowers early in the year before the fronds develop fully. It is the food plant for the

weevil Weevils are beetles belonging to the Taxonomic rank, superfamily Curculionoidea, known for their elongated snouts. They are usually small, less than in length, and Herbivore, herbivorous. Approximately 97,000 species of weevils are known. They b ...

, ''

Procas granulicollis ''Procas granulicollis'' (sometimes known as the climbing corydalis weevil) is a beetle in the family Curculionidae, the true weevils. It was first described in 1848 from a specimen collected in Cumbria and has since been found at a number of oth ...

'' and the

beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...

, ''

Sirocalodes mixtus ''Sirocalodes'' is a genus of beetles belonging to the family Curculionidae The Curculionidae are a family of weevils, commonly called snout beetles or true weevils. They are one of the largest animal families, with 6,800 genera and 83,000 spe ...

''. Pollination is by

honey bees A honey bee (also spelled honeybee) is a eusocial flying insect within the genus ''Apis'' of the bee clade, all native to Afro-Eurasia. After bees spread naturally throughout Africa and Eurasia, humans became responsible for the current cosmo ...

and

bumblebee A bumblebee (or bumble bee, bumble-bee, or humble-bee) is any of over 250 species in the genus ''Bombus'', part of Apidae, one of the bee families. This genus is the only extant group in the tribe Bombini, though a few extinct related gener ...

s. erspectives in Plant Ecology, Evolution and Systematics - Biological Flora of Central Europe: Ceratocapnos claviculata/ref>

References

BSBI Species Accounts - Ceratocapnos claviculataPerspectives in Plant Ecology, Evolution and Systematics - Biological Flora of Central Europe: Ceratocapnos claviculata
{{Taxonbar, from=Q160220 Fumarioideae Plants described in 1984 Flora of Europe Flora of Belgium Flora of Denmark Flora of Germany Flora of Ireland Flora of the Netherlands Flora of Spain Flora of Portugal Flora of Norway Flora of the United Kingdom Taxa named by Magnus Lidén