
The cephalothorax, also called prosoma in some groups, is a
tagma of various
arthropod
Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s, comprising the head and the
thorax
The thorax (: thoraces or thoraxes) or chest is a part of the anatomy of mammals and other tetrapod animals located between the neck and the abdomen.
In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main di ...
fused together, as distinct from the
abdomen
The abdomen (colloquially called the gut, belly, tummy, midriff, tucky, or stomach) is the front part of the torso between the thorax (chest) and pelvis in humans and in other vertebrates. The area occupied by the abdomen is called the abdominal ...
behind.
(The terms ''prosoma'' and ''opisthosoma'' are equivalent to ''cephalothorax'' and ''abdomen'' in some groups. The terms ''prosoma'' and ''opisthosoma'' may be preferred by some researchers in cases such as arachnids, where there is neither fossil nor embryonic evidence animals in this class have ever had separate heads and thoraxes, and where the ''opisthosoma'' contains organs atypical of a true ''abdomen'', such as a heart and respiratory organs.) The word ''cephalothorax'' is derived from the
Greek words for head (, ') and thorax (, '). This fusion of the head and thorax is seen in
chelicerates and
crustaceans; in other groups, such as the
Hexapoda (including
insect
Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s), the head remains free of the thorax.
In
horseshoe crabs and many
crustaceans, a hard shell called the
carapace covers the cephalothorax.
Arachnid anatomy
Fovea
The fovea is the centre of the cephalothorax and is located behind the head (only in spiders).
[Dalton, Steve (2008). ''Spiders; The Ultimate Predators''. A & C Black, London. P.p. 19. .] It is often important in identification. It can be transverse or procurved
[Smith, A. M. (1990c). Baboon spiders: Tarantulas of Africa and the Middle East. Fitzgerald Publishing, London, pp. 138. Retrieved February 13, 2016.] and can, in some
tarantulas (e.g. ''
Ceratogyrus darlingi'') have a "horn".
Clypeus
The clypeus is the space between the anterior of the cephalothorax and the ocularium. It is found in most arachnids.
[ It is connected to the labrum of the invertebrate, between the labrum and the face.
]
Ocularium
The ocularium is a "turret" for the ocelli found in most arachnids.[Spiders...]
Yorkshire Naturalists' Union. Retrieved February 13, 2016. In harvestmen, it may have the ornament of spines.[Sankey, John & Savory, Theodore. ''British Harvestmen''. Academic Press. P.p. 1–75. .]
Trident
The trident is a small group of (usually three) spines found in harvestmen exclusively. It is located in front of the ocularium. It varies in size amongst species; in some it is completely absent, and in others it is enlarged considerably.[
]
References
Arthropod anatomy
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