Cephalopholis Boenak
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''Cephalopholis boenak'', the chocolate hind, brownbarred rockcod, brown-banded cod or brown-banded rockcod, is a species of marine
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...
, a
grouper Groupers are fish of any of a number of genera in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes. Not all serranids are called "groupers"; the family also includes the sea basses. The common name "grouper" is u ...
from the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Epinephelinae Groupers are fish of any of a number of genera in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes. Not all serranids are called "groupers"; the family also includes the sea basses. The common name "grouper" is u ...
which is in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Serranidae The Serranidae are a large family of fishes belonging to the order Perciformes. The family contains about 450 species in 65 genera, including the sea basses and the groupers (subfamily Epinephelinae). Although many species are small, in some case ...
which also includes the
anthias Anthias are members of the family Serranidae and make up the subfamily Anthiinae. Anthias make up a sizeable portion of the population of pink, orange, and yellow reef fishes seen swarming in most coral reef photography and film. The name Anthi ...
and sea basses. It is associated with reefs over a wide
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
distribution. It is an important species for commercial fisheries in some parts of its range.


Description

''Cephalopholis boenak'' has a body which has a
standard length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish m ...
which is 2.6–3.1 times its depth and with a head that is around a fifth of the standard length. Its
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...
contains 9 spines and 15–17 soft rays while the
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
has 3 spines and 8 soft rays. The preopercle is rounded with fine serrations but no enlarged spines at its angle and with a flesh lower margin. The body, including the abdomen, is covered in
ctenoid A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scales, which can also provide effective camouflage through the use of reflection and colouration, as ...
scales which is a characteristic of this species. The overall colour is brownish to greenish-grey and there are normally 7–8 dark vertical bars on the flanks. The dorsal fin has a wide dark submarginal band and a thin white margin, these are also present on the anal fins, and the upper and lower lobes of the
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
. There is a blackish spot on the upper margin of the gill cover. The juveniles have a yellowish on the posterior portion of their body and are marked with a white stripe which runs from the snout to the origin of the dorsal fin. It grows to a maximum
total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...
of .


Distribution

''Cephalopholis boenak'' has a wide distribution in the Indian Ocean and the
Western Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
from the eastern coast of Africa where it occurs from Kenya south to southern Mozambique along the southern Asian coast and into the Pacific where it occurs north as far as the
Ryukyu Islands The , also known as the or the , are a chain of Japanese islands that stretch southwest from Kyushu to Taiwan: the Ōsumi, Tokara, Amami, Okinawa, and Sakishima Islands (further divided into the Miyako and Yaeyama Islands), with Yonaguni ...
and east to
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
. In Australia it has been recorded along the northern coast from
Coral Bay, Western Australia Coral Bay is a small coastal settlement located north of Perth, in the Shire of Carnarvon in the Gascoyne region of Western Australia. Bordered by the Ningaloo Reef, it is a popular tourist destination and largely owes its survival to revenue ...
to
Moreton Bay Moreton Bay is a bay located on the eastern coast of Australia from central Brisbane, Queensland. It is one of Queensland's most important coastal resources. The waters of Moreton Bay are a popular destination for recreational anglers and are ...
in
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...
.


Habitat and biology

''Cephalopholis boenak'' is mainly a coastal species which is found on silty, dead and living coral reefs as deep as . It is a carnivore which preys mainly on other fishes and on
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
s. The chocolate hind is a
protogynous hermaphrodite Sequential hermaphroditism (called dichogamy in botany) is a type of hermaphroditism that occurs in many fish, gastropods, and plants. Sequential hermaphroditism occurs when the individual changes its sex at some point in its life. In particular, ...
, they undergo a sex change from females to males, although some are born as males. They occur in relatively small social groups which comprise a single male, one or two smaller females, and a variable number of sexually inactive individuals. These sexually inactive fish are sexually inactive adults or juveniles. The males defend a
territory A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...
from other males and the territory is much larger than the home ranges of the females. The males visit the females and other fish within their territory at specific locations, almost always using the same route. Courtship occurs in the late afternoons but breeding behaviour is not associated with any particular moon phase and can happen multiple times during the breeding season.


Utilisation

''Cephalopholis boenak'' is commercially fished for human consumption in many parts of its range. Hook-and-line, traps and trawling are the main methods used to catch them, although they are occasionally caught using gillnet and purse-seine. In
Hong Kong Hong Kong ( (US) or (UK); , ), officially the Hong Kong Special Administrative Region of the People's Republic of China ( abbr. Hong Kong SAR or HKSAR), is a city and special administrative region of China on the eastern Pearl River Delt ...
it has increased in commercial importance as a result of the decline of larger-bodied species of grouper and the chocolate hind is now one of the more important species in the Hong Kong fish market It is commonly landed by artisanal fishermen in southern Kenya. In Sri Lamka it is also harvested to be traded in the marine
aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...
trade.


Taxonomy

''Cephalopholis boenak'' was first formally described as ''Bodianus boenak'' by the
German German(s) may refer to: * Germany (of or related to) **Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ger ...
naturalist
Marcus Elieser Bloch Marcus Elieser Bloch (1723–1799) was a German physician and naturalist who is best known for his contribution to ichthyology through his multi-volume catalog of plates illustrating the fishes of the world. Brought up in a Hebrew-speaking Jewish ...
(1723–1799) with the type locality given erroneously as Japan when it was actually Indonesia.


References


External links

* {{Taxonbar, from=Q2791901 boenak Fish of the Indian Ocean Fish of the Pacific Ocean Fish of Thailand Fish described in 1790 Taxa named by Marcus Elieser Bloch Taxonomy articles created by Polbot