Central Asian Red Deer
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The Central Asian red deer (''Cervus hanglu''), also known as the Tarim red deer is a
deer Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindeer ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
native to
Central Asia Central Asia, also known as Middle Asia, is a subregion, region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes t ...
, where it used to be widely distributed, but is scattered today with small population units in several countries. It has been listed as
Least Concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
since 2017. It was first described in the mid-19th century.


Characteristics

The Central Asian red deer's fur is light ginger in colour.


Taxonomy

The
scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Cervus hanglu'' was proposed by
Johann Andreas Wagner Johann Andreas Wagner (21 March 1797 – 17 December 1861) was a German palaeontologist, zoologist and archaeologist who wrote several important works on palaeontology. Career Wagner was a professor at the University of Munich, and curator ...
in 1844 for a deer
specimen Specimen may refer to: Science and technology * Sample (material), a limited quantity of something which is intended to be similar to and represent a larger amount * Biological specimen or biospecimen, an organic specimen held by a biorepository ...
from
Kashmir Kashmir () is the northernmost geographical region of the Indian subcontinent. Until the mid-19th century, the term "Kashmir" denoted only the Kashmir Valley between the Great Himalayas and the Pir Panjal Range. Today, the term encompas ...
that differed from the
red deer The red deer (''Cervus elaphus'') is one of the largest deer species. A male red deer is called a stag or hart, and a female is called a hind. The red deer inhabits most of Europe, the Caucasus Mountains region, Anatolia, Iran, and parts of wes ...
(''Cervus elaphus'') in the shape and points of the
antler Antlers are extensions of an animal's skull found in members of the Cervidae (deer) family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally found only on male ...
s. In the 19th and early 20th centuries, the following red deer specimens from Central Asia were described: *''Cervus cashmeriensis'' was proposed by
Andrew Leith Adams Andrew Leith Adams FRSE, FRS (21 March 1827 – 29 July 1882) was a Scottish physician, naturalist and geologist. He was the father of the writer Francis Adams.Gaston, A. J. in Oxford Dictionary of National Biography. Volume 1. pp. 222–223 ...
in 1858 for the red deer occurring in the montane forests of Kashmir. *''Cervus yarkandensis'' proposed by
William Thomas Blanford William Thomas Blanford (7 October 183223 June 1905) was an English geologist and naturalist. He is best remembered as the editor of a major series on ''The Fauna of British India, Including Ceylon and Burma''. Biography Blanford was born ...
in 1892 was a red deer stag killed in the
Tarim River The Tarim River ( zh, p=Tǎlǐmù Hé, c=塔里木河; ug, تارىم دەرياسى, Tarim deryasi), known in Sanskrit as the Śītā, is an endorheic river in Xinjiang, China. It is the principal river of the Tarim Basin, a desert region of C ...
basin,
East Turkestan East Turkestan ( ug, شەرقىي تۈركىستان, Sherqiy Türkistan, bold=no; zh, s=东突厥斯坦; also spelled East Turkistan), is a loosely-defined geographical and historical region in the western provinces of the People's Republic of ...
. *''Cervus bactrianus'' proposed by
Richard Lydekker Richard Lydekker (; 25 July 1849 – 16 April 1915) was an English naturalist, geologist and writer of numerous books on natural history. Biography Richard Lydekker was born at Tavistock Square in London. His father was Gerard Wolfe Lydekker, ...
in 1900 was a live deer caught in the vicinity of
Tashkent Tashkent (, uz, Toshkent, Тошкент/, ) (from russian: Ташкент), or Toshkent (; ), also historically known as Chach is the capital and largest city of Uzbekistan. It is the most populous city in Central Asia, with a population of ...
in
Uzbekistan Uzbekistan (, ; uz, Ozbekiston, italic=yes / , ; russian: Узбекистан), officially the Republic of Uzbekistan ( uz, Ozbekiston Respublikasi, italic=yes / ; russian: Республика Узбекистан), is a doubly landlocked cou ...
and brought to
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
. Two years later, he considered this ungulate to be a red deer
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
(''C. e. bactrianus''). *''Cervus hagenbeckii'' proposed by a Russian zoologist in 1904 for a red deer from
Russian Turkestan Russian Turkestan (russian: Русский Туркестан, Russkiy Turkestan) was the western part of Turkestan within the Russian Empire’s Central Asian territories, and was administered as a Krai or Governor-Generalship. It comprised the ...
that was sent to the
Moscow Zoo The Moscow Zoo or Moskovsky Zoopark (russian: Московский зоопарк) is a zoo founded in 1864 by professor-biologists, K.F. Rulje, S.A. Usov and A.P. Bogdanov, from the Moscow State University. In 1919, the zoo was nationalized. In ...
in the 1890s. In 1951, John Ellerman and
Terence Morrison-Scott Sir Terence Charles Stuart Morrison-Scott (24 October 1908 – 25 November 1991) was a British zoologist who was Director of the Science Museum and the British Museum (Natural History) in London, England.CranbrookScott, Sir Terence Charles Stu ...
recognised all these specimens as subspecies of the red deer. In 2005, Peter Grubb also considered the proposed
taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
as subspecies of the red deer. IUCN Red List assessors provisionally recognised its status as a distinct species in 2017; The Central Asian red deer is thought to comprise three subspecies: *the
Kashmir stag The Kashmir stag (''Cervus hanglu hanglu''), also called hangul (), is a subspecies of Central Asian red deer endemic to Kashmir and surrounding areas. It is found in dense riverine forests in the high valleys and mountains of Jammu and Kashmi ...
(''C. h. hanglu''), the
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
, is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to Kashmir in
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
; *the
Bactrian deer The Bactrian deer (''Cervus hanglu bactrianus''), also called the Bukhara deer, Bokhara deer, or Bactrian wapiti, is a lowland subspecies of Central Asian red deer native to Central Asia. It is similar in ecology to the related Yarkand deer (''C ...
(''C. h. bactrianus'') ranges throughout Central Asia; *the
Yarkand deer The Yarkand deer (''Cervus hanglu yarkandensis''), also known as the Theenivs deer, Tarim deer, or Lop Nor stag, is a subspecies of the Central Asian red deer that is native to the province of Xinjiang, China. It is similar in ecology to the rel ...
(''C. h. yarkandensis'') ranges in
Xinjiang Xinjiang, SASM/GNC: ''Xinjang''; zh, c=, p=Xīnjiāng; formerly romanized as Sinkiang (, ), officially the Xinjiang Uygur Autonomous Region (XUAR), is an autonomous region of the People's Republic of China (PRC), located in the northwest ...
Province of
western China Western China (, or rarely ) is the west of China. In the definition of the Chinese government, Western China covers one municipality (Chongqing), six provinces (Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively r ...
.


Phylogeny

An analysis of
mitochondrial DNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial D ...
of 125 tissue samples from 50 populations of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Cervus'' included two samples from
Tajikistan Tajikistan (, ; tg, Тоҷикистон, Tojikiston; russian: Таджикистан, Tadzhikistan), officially the Republic of Tajikistan ( tg, Ҷумҳурии Тоҷикистон, Jumhurii Tojikiston), is a landlocked country in Centr ...
and three from western China. The results supported the classification of the red deer populations in Central Asia as two distinct red deer subspecies. Results of a subsequent
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
analysis of
Cervinae The Cervinae or the Old World deer, are a subfamily of deer. Alternatively, they are known as the plesiometacarpal deer, due to their ankle structure being different from the telemetacarpal deer of the Capreolinae. Classification and species The ...
tissue samples indicated that deer samples from Central Asia form a distinct
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
and warrant to be raised to
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
level. The Central Asian red deer group appears to have genetically diverged from the European red deer group during the
Chibanian The Chibanian, widely known by its previous designation of Middle Pleistocene, is an Age (geology), age in the international geologic timescale or a Stage (stratigraphy), stage in chronostratigraphy, being a division of the Pleistocene Epoch withi ...
period between 770,000 and 126,000 years ago. The first phylogenetic analysis using hair samples of the deer population in
Dachigam National Park Dachigam National Park is a national park located, from Srinagar city in Srinagar district of Jammu and Kashmir, India on the east side of Dal Lake. It covers an area of 161, 62.16(updated recently) by hgtian student SIJ. The name of the park ...
in Jammu and Kashmir was published in 2015. Results showed that these samples form a sub
cluster may refer to: Science and technology Astronomy * Cluster (spacecraft), constellation of four European Space Agency spacecraft * Asteroid cluster, a small asteroid family * Cluster II (spacecraft), a European Space Agency mission to study t ...
within the Central Asian red deer group; they are genetically closer to this group than to the European red deer.


References


External links

* {{Taxonbar, from=Q5060377 Mammals of Asia