''Celtis reticulata'', with common names including netleaf hackberry, western hackberry, Douglas hackberry,DeBolt, Ann M. (2002
"''Celtis reticulata'' Torr. netleaf hackberry"
United States Forest Service netleaf sugar hackberry, palo blanco, and acibuche, is a small- to medium-sized

deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

native to western

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

."Netleaf Hackberry"
Tree New Mexicohttp://plants.usda.gov/java/profile?symbol=CELAR&mapType=large&photoID=celar_001_ahp.tif USDA: profile

Description

''Celtis reticulata'' usually grows to a small-sized tree, in height and mature at in diameter, although some individuals are known up to high and thick. It is often scraggly, stunted or even a large bush.
United States Forest Service It grows at altitudes of .http://ucjeps.berkeley.edu/cgi-bin/get_JM_treatment.pl?7729,7730,7731 Jepson Hackberry

bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, ...

is gray to brownish gray with the trunk bark forming vertical corky ridges that are checkered between the furrows. The young twigs are

puberulent Trichomes (); ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a plant ...

, or covered with very fine hairs. The blade of the

leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...

can be long, usually about . They are lanceolate to ovate, disproportionate at the base, leathery, entire to serrate (tending toward serrate), clearly net-veined, base obtuse to more or less cordate, tip obtuse to acuminate, and scabrous, with a dark green upper surface and a yellowish-green lower surface. The small stalks attaching the leaf blade to the stem (the petioles) are generally about long. The flowers are very small, averaging 2 mm across. They form singly, or in cymose clustersBenson, Lyman D. and Darrow, Robert A. (1981) "Celtis: Hackberry, Palo Blanco" ''Trees and Shrubs of the Southwestern Deserts'' (3rd edition) University of Arizona Press, Tucson, Arizona, pages 154-155 pedicel in fr 4–15 mm. The fruit is a rigid, brownish to purple berry, 5 to 12 mm in diameter, with thin, sweet pulp. If uneaten, they can stay on the plant through early winter.

Similar species

''C. reticulata'' is often confused with the related species ''

Celtis pallida ''Celtis ehrenbergiana'', called the desert hackberry or spiny hackberry, is a plant species that has long been called ''C. pallida'' by many authors, including in the "Flora of North America" database. It is native to Arizona, Florida, New Mexic ...

'', the spiny hackberry or desert hackberry, ''

Celtis occidentalis ''Celtis occidentalis'', commonly known as the common hackberry, is a large deciduous tree native to North America. It is also known as the nettletree, sugarberry, beaverwood, northern hackberry, and American hackberry. It is a moderately long-l ...

'', the common hackberry, and ''

Celtis laevigata ''Celtis laevigata'' is a medium-sized tree native to North America. Common names include sugarberry, Southern hackberry, or in the southern U.S. sugar hackberry or just hackberry. Sugarberry is easily confused with common hackberry ('' C. occ ...

'', the sugarberry or southern hackberry.

Distribution and habitat

Prehistoric

''Celtis reticulata'' was one of the species analyzed in a

pollen core A pollen core is a core sample of a medium containing a stratigraphic sequence of pollen. Analysis of the type and frequency of the pollen in each layer is used to study changes in climate or land use using regional vegetation as a proxy. This a ...

sampling study in northern

Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...

, in which the early to late

Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togethe ...

flora Flora is all the plant life present in a particular region or time, generally the naturally occurring (indigenous) native plants. Sometimes bacteria and fungi are also referred to as flora, as in the terms '' gut flora'' or '' skin flora''. E ...

association was reconstructed; this study in the

Waterman Mountains The Waterman Mountains are a low mountainous landform in Pima County, United States. Notable among the tree species is the elephant tree (''Bursera microphylla'') which species exhibits a contorted multi-furcate architecture; most of these froze i ...

(

Pima County, Arizona Pima County ( ) is a county in the south central region of the U.S. state of Arizona. As of the 2020 census, the population was 1,043,433, making it Arizona's second-most populous county. The county seat is Tucson, where most of the population ...

) demonstrated that ''C. reticulata'' was found to be present after the

Wisconsinan glaciation The Wisconsin Glacial Episode, also called the Wisconsin glaciation, was the most recent glacial period of the North American ice sheet complex. This advance included the Cordilleran Ice Sheet, which nucleated in the northern North American Cor ...

, but is not a current

taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...

of this former

Pinyon–juniper woodland Pinyon–juniper woodland, also spelled piñon–juniper woodland, is a vegetation type (biome) of Western United States higher elevation deserts, characterized by being an open forest dominated by low, bushy, evergreen junipers (''Juniperus oste ...

area which is now in central and northern Arizona.

Current

At its western edge, the tree's natural range includes the

Columbia River Basin The Columbia River drainage basin is the drainage basin of the Columbia River in the Pacific Northwest region of North America. It covers . In common usage, the term often refers to a smaller area, generally the portion of the drainage basin that ...

Oregon Oregon () is a U.S. state, state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington (state), Washington, while the Snake River delineates much of it ...

Washington Washington commonly refers to: * Washington (state), United States * Washington, D.C., the capital of the United States ** A metonym for the federal government of the United States ** Washington metropolitan area, the metropolitan area centered on ...

, and western

Idaho Idaho ( ) is a state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Montana and Wyom ...

. It can also be found in

Southern California Southern California (commonly shortened to SoCal) is a geographic and Cultural area, cultural region that generally comprises the southern portion of the U.S. state of California. It includes the Los Angeles metropolitan area, the second most po ...

in the southwestern

Sierra Nevada The Sierra Nevada () is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primarily ...

foothills, the

Peninsular Ranges The Peninsular Ranges (also called the Lower California province) are a group of mountain ranges that stretch from Southern California to the southern tip of the Baja California peninsula; they are part of the North American Coast Ranges, which ...

and eastern

Transverse Ranges The Transverse Ranges are a group of mountain ranges of southern California, in the Pacific Coast Ranges physiographic region in North America. The Transverse Ranges begin at the southern end of the California Coast Ranges and lie within Santa B ...

, and the

Mojave Desert The Mojave Desert ( ; mov, Hayikwiir Mat'aar; es, Desierto de Mojave) is a desert in the rain shadow of the Sierra Nevada mountains in the Southwestern United States. It is named for the indigenous Mojave people. It is located primarily in ...

sky island Sky islands are topographic isolation, isolated mountains surrounded by radically different lowland environments. The term originally referred to those found on the Mexican Plateau, and has extended to similarly isolated montane ecosystems, hi ...

s. Its central range includes the

Rio Grande The Rio Grande ( and ), known in Mexico as the Río Bravo del Norte or simply the Río Bravo, is one of the principal rivers (along with the Colorado River) in the southwestern United States and in northern Mexico. The length of the Rio G ...

watershed and the

Chihuahuan Desert The Chihuahuan Desert ( es, Desierto de Chihuahua, ) is a desert ecoregion designation covering parts of northern Mexico and the southwestern United States. It occupies much of far West Texas, the middle to lower Rio Grande Valley and the lower P ...

in southern

and

New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...

, western Texas, and northern

Sonora Sonora (), officially Estado Libre y Soberano de Sonora ( en, Free and Sovereign State of Sonora), is one of the 31 states which, along with Mexico City, comprise the Administrative divisions of Mexico, Federal Entities of Mexico. The state is d ...

Chihuahua Chihuahua may refer to: Places *Chihuahua (state), a Mexican state **Chihuahua (dog), a breed of dog named after the state **Chihuahua cheese, a type of cheese originating in the state **Chihuahua City, the capital city of the state **Chihuahua Mun ...

Coahuila Coahuila (), formally Coahuila de Zaragoza (), officially the Free and Sovereign State of Coahuila de Zaragoza ( es, Estado Libre y Soberano de Coahuila de Zaragoza), is one of the 32 states of Mexico. Coahuila borders the Mexican states of N ...

. It is also found in the

Madrean Sky Islands The Madrean Sky Islands are enclaves of Madrean pine–oak woodlands, found at higher elevations in a complex of small mountain ranges in southern and southeastern Arizona, southwestern New Mexico, and northwestern Mexico. The sky islands are sur ...

of the

Sierra Madre Occidental The Sierra Madre Occidental is a major mountain range system of the North American Cordillera, that runs northwest–southeast through northwestern and western Mexico, and along the Gulf of California. The Sierra Madre is part of the American C ...

in northern Sonora, and in the White Mountains and along the

Mogollon Rim The Mogollon Rim ( or or ) is a topographical and geological feature cutting across the northern half of the U.S. state of Arizona. It extends approximately , starting in northern Yavapai County and running eastward, ending near the border ...

in Arizona. The banks of the

Colorado River The Colorado River ( es, Río Colorado) is one of the principal rivers (along with the Rio Grande) in the Southwestern United States and northern Mexico. The river drains an expansive, arid drainage basin, watershed that encompasses parts of ...

also provide suitable habitat, from the

Grand Canyon The Grand Canyon (, yuf-x-yav, Wi:kaʼi:la, , Southern Paiute language: Paxa’uipi, ) is a steep-sided canyon carved by the Colorado River in Arizona, United States. The Grand Canyon is long, up to wide and attains a depth of over a m ...

northeast through

Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...

to western

Colorado Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of t ...

. Its easternmost natural range is in the hills of

Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...

Oklahoma Oklahoma (; Choctaw language, Choctaw: ; chr, ᎣᎧᎳᎰᎹ, ''Okalahoma'' ) is a U.S. state, state in the South Central United States, South Central region of the United States, bordered by Texas on the south and west, Kansas on the nor ...

Kansas Kansas () is a state in the Midwestern United States. Its capital is Topeka, and its largest city is Wichita. Kansas is a landlocked state bordered by Nebraska to the north; Missouri to the east; Oklahoma to the south; and Colorado to the ...

, and

Louisiana Louisiana , group=pronunciation (French: ''La Louisiane'') is a state in the Deep South and South Central regions of the United States. It is the 20th-smallest by area and the 25th most populous of the 50 U.S. states. Louisiana is borde ...

. The species grows in

alluvial soils Alluvium (from Latin ''alluvius'', from ''alluere'' 'to wash against') is loose clay, silt, sand, or gravel that has been deposited by running water in a stream bed, on a floodplain, in an alluvial fan or beach, or in similar settings. Allu ...

and rocky sites far above the water line. It is very drought tolerant, accepting sites with only in annual precipitation.

Ecology

The leaves are eaten by a number of insects, particularly certain

moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...

caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...

s. The berries are eaten by wildlife, including birds.

Mule deer The mule deer (''Odocoileus hemionus'') is a deer indigenous to western North America; it is named for its ears, which are large like those of the mule. Two subspecies of mule deer are grouped into the black-tailed deer. Unlike the related whit ...

and

bighorn sheep The bighorn sheep (''Ovis canadensis'') is a species of sheep native to North America. It is named for its large horns. A pair of horns might weigh up to ; the sheep typically weigh up to . Recent genetic testing indicates three distinct subspec ...

eat the fresh twigs. Beavers feed on the plant as well.

Cultivation

''Celtis reticulata'' is cultivated by

plant nurseries A nursery is a place where plants are propagated and grown to a desired size. Mostly the plants concerned are for gardening, forestry or conservation biology, rather than agriculture. They include retail nurseries, which sell to the general p ...

and available as an

ornamental plant Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that i ...

for

native plant In biogeography, a native species is indigenous to a given region or ecosystem if its presence in that region is the result of only local natural evolution (though often popularised as "with no human intervention") during history. The term is equ ...

drought-tolerant Drought tolerance is the ability to which a plant maintains its biomass production during arid or drought conditions. Some plants are naturally adapted to dry conditions'','' surviving with protection mechanisms such as desiccation tolerance, detox ...

natural landscape A natural landscape is the original landscape that exists before it is acted upon by human culture Culture () is an umbrella term which encompasses the social behavior, institutions, and norms found in human societies, as well as the kn ...

, and

habitat garden A wildlife garden (or wild garden) is an environment created by a gardener that serves as a sustainable haven for surrounding wildlife. Wildlife gardens contain a variety of habitats that cater to native and local plants, birds, amphibians, ...

s, and for ecological restoration projects.

Uses

The berries and seeds have long been used as a food source by Native Americans of the

Southwestern United States The Southwestern United States, also known as the American Southwest or simply the Southwest, is a geographic and cultural region of the United States that generally includes Arizona, New Mexico, and adjacent portions of California, Colorado, Ne ...

, including the

Apache The Apache () are a group of culturally related Native American tribes in the Southwestern United States, which include the Chiricahua, Jicarilla, Lipan, Mescalero, Mimbreño, Ndendahe (Bedonkohe or Mogollon and Nednhi or Carrizaleño an ...

(

Chiricahua Chiricahua ( ) is a band of Apache Native Americans. Based in the Southern Plains and Southwestern United States, the Chiricahua (Tsokanende ) are related to other Apache groups: Ndendahe (Mogollon, Carrizaleño), Tchihende (Mimbreño), Sehende ...

and

Mescalero Mescalero or Mescalero Apache ( apm, Naa'dahéńdé) is an Apache tribe of Southern Athabaskan–speaking Native Americans. The tribe is federally recognized as the Mescalero Apache Tribe of the Mescalero Apache Reservation, located in south-cen ...

), both fresh and preserved, and the

Navajo The Navajo (; British English: Navaho; nv, Diné or ') are a Native American people of the Southwestern United States. With more than 399,494 enrolled tribal members , the Navajo Nation is the largest federally recognized tribe in the United ...

, who eat them both fresh and ground.

References

External links

Jepson Manual treatment for ''Celtis reticulata''Celtis reticulata Range MapLady Bird Johnson database & gallery: Celtis reticulataUniv. of Wash. Burke Museum: photo gallery''Celtis reticulata'' - CalPhotos gallery
{{Taxonbar, from=Q4215121 reticulata Trees of the Northwestern United States Trees of the Southwestern United States Trees of the Plains-Midwest (United States) Flora of Arizona Flora of California Flora of New Mexico Flora of the Cascade Range Flora of the Chihuahuan Desert Flora of the Rocky Mountains Flora of the Sierra Nevada (United States) Flora of the Sonoran Deserts Natural history of the California chaparral and woodlands Natural history of the Mojave Desert Natural history of the Peninsular Ranges Natural history of the Transverse Ranges Plants described in 1824 Taxa named by John Torrey Bird food plants Butterfly food plants Plants used in Native American cuisine Garden plants of North America Drought-tolerant trees Ornamental trees Flora without expected TNC conservation status