Celmisia Spectabilis
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''Celmisia spectabilis'', also known as cotton daisy or by its Māori name puharetaiko, is a mountain daisy in the family
Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae w ...
, and is endemic to New Zealand, where it is one of the most widespread species in the genus ''
Celmisia ''Celmisia'' (New Zealand aster or New Zealand daisy) is a genus of perennial herbs or subshrubs, in the family Asteraceae. Most of the species are endemic to New Zealand; several others are endemic to Australia. ; Species and nothospecies #RED ...
''.


Taxonomy

Joseph Hooker described this species in 1844, in the first volume of his ''Flora Antarctica''. The specimens he studied were collected by the English botanist John Bidwill in 1839 on
Mt Tongariro Mount Tongariro (; ) is a compound volcano in the Taupō Volcanic Zone of the North Island of New Zealand. It is located to the southwest of Lake Taupō, and is the northernmost of the three active volcanoes that dominate the landscape of th ...
, on New Zealand's North Island.


Description

''C. spectabilis'' has is a robust plant with leathery leaves that are
ovate Ovate may refer to: *Ovate (egg-shaped) leaves, tepals, or other botanical parts *Ovate, a type of prehistoric stone hand axe *Ovates, one of three ranks of membership in the Welsh Gorsedd *Vates In modern English, the nouns vates () and ovat ...
to
lanceolate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...
or narrowly
oblong An oblong is a non-square rectangle. Oblong may also refer to: Places * Oblong, Illinois, a village in the United States * Oblong Township, Crawford County, Illinois, United States * A strip of land on the New York-Connecticut border in the Unit ...
, and are usually 3–18 cm long by 1–2 cm wide. They have a shiny, green upper surface, usually close to hairless except in young or North Island plants. The leaf underside is densely covered in soft, whitish or buff-coloured felted woolly hairs. The leaf margins usually roll downwards. The leaf bases overlap and compact to form a stout
pseudostem A stem is one of two main structural axes of a vascular plant, the other being the root. It supports leaves, flowers and fruits, transports water and dissolved substances between the roots and the shoots in the xylem and phloem, stores nutrien ...
, and slowly rot away forming a damp mass that can protect young leaves from grass fires. ''C. spectabilis'' can grow as a single rosette up to 1 m wide, or several, forming mats up to 2 m across. The flower stems reach 30 cm tall and are densely covered with loose white hairs. A showy solitary flower head, 3–5 cm across, is borne at the end of each stem. The numerous ray florets are white and the disc florets yellow. This species hybridises with '' C. lyallii'' to form ''C''. × ''pseudolyallii'', which resembles ''C. spectabilis'' but with longer narrower leaves. File:Celmisia spectabilis 305355591.jpg, Flower File:Celmisia spectabilis spectabilis 300695981.jpg, Bracts File:Celmisia spectabilis spectabilis 341005633.jpg, Leaf underside and flower stalks File:Celmisia spectabilis spectabilis 346115196.jpg, Growing as a clump


Distribution

This species is found in montane to low-alpine regions (300–1700) through the both main islands, usually in the east. In the
North Island The North Island, also officially named Te Ika-a-Māui, is one of the two main islands of New Zealand, separated from the larger but much less populous South Island by the Cook Strait. The island's area is , making it the world's 14th-largest ...
it is found south of the
Volcanic Plateau A volcanic plateau is a plateau produced by volcanic activity. There are two main types: lava plateaus and pyroclastic plateaus. Lava plateau Lava plateaus are formed by highly fluid basaltic lava during numerous successive eruptions throu ...
and Mount Hikurangi, in tussock and fellfields, and in snow tussock in the Tararua Range (but is not found in the eastern Wairarapa). In the
South Island The South Island, also officially named , is the larger of the two major islands of New Zealand in surface area, the other being the smaller but more populous North Island. It is bordered to the north by Cook Strait, to the west by the Tasman ...
is occurs in higher-rainfall habitats from north-west Nelson to Arthur's Pass; Nelson plants are smaller and less robust than those from
Canterbury Canterbury (, ) is a City status in the United Kingdom, cathedral city and UNESCO World Heritage Site, situated in the heart of the City of Canterbury local government district of Kent, England. It lies on the River Stour, Kent, River Stour. ...
. The Nelson plants were previously classified as their own variety, ''Celmisia spectabilis'' var. ''angustifolia'', but leaf size increases gradually with latitude and there is no clear distinction.


Ecology

''Celmisia spectabilis'' flowers in early to mid summer, and has many pollinators, including ''
Melangyna novaezelandiae ''Melangyna novaezelandiae'' (commonly referred to as the "large hoverfly") is a hoverfly endemic to New Zealand.Parkinson, Brian (2007). ''A Photographic Guide To The Insects Of New Zealand''. New Holland Publishers (NZ) Ltd. . It is a generaliz ...
'' and ''
Leioproctus ''Leioproctus'' is a genus in the plaster bee family Colletidae. Its members are primarily found in Australasia and temperate South America, and include the most common native bees in New Zealand. Species It includes the following species: *' ...
.''


Conservation

Because it was able to withstand once-frequent tussock burning, and is unpalatable to
sheep Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus ''Ovis'', in everyday usage it almost always refers to domesticated s ...
, this daisy is now one of the more widespread species in the mountainous areas of New Zealand. One subspecies, ''Celmisia spectabilis lanceolata'', is considered "At Risk—Naturally Uncommon" in the
Department of Conservation An environmental ministry is a national or subnational government agency politically responsible for the environment and/or natural resources. Various other names are commonly used to identify such agencies, such as Ministry of the Environment ...
NZTCS The New Zealand Threat Classification System is used by the Department of Conservation to assess conservation priorities of species in New Zealand. The system was developed because the IUCN Red List, a similar conservation status system, had some ...
database.


References


External links

*
iNaturalist observations
of ''Celmisia spectabilis'' * * ''Celmisia spectabilis'' a

{{Taxonbar, from=Q2944119 spectabilis Flora of New Zealand Plants described in 1844 Taxa named by Joseph Dalton Hooker