Cedarosaurus Parts
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''Cedarosaurus'' (meaning "Cedar lizard" - named after the
Cedar Mountain Formation The Cedar Mountain Formation is the name given to a distinctive sedimentary geologic formation in eastern Utah, spanning most of the early and mid- Cretaceous. The formation was named for Cedar Mountain in northern Emery County, Utah, where Will ...
, in which it was discovered) was a nasal-crested
macronaria Macronaria is a clade of sauropod dinosaurs. Macronarians are named after the large diameter of the nasal opening of their skull, known as the external naris, which exceeded the size of the orbit, the skull opening where the eye is located (hence ...
n
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
from the Early
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
Period (
Valanginian In the geologic timescale, the Valanginian is an age or stage of the Early or Lower Cretaceous. It spans between 139.8 ± 3.0 Ma and 132.9 ± 2.0 Ma (million years ago). The Valanginian Stage succeeds the Berriasian Stage of the Lower Cretace ...
). It was a sauropod which lived in what is now
Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...
. The fossils were discovered in 1996 in eastern
Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...
within the rocks of the Yellow Cat Member of the Cedar Mountain Formation. It was officially named and described by Tidwell, Carpenter and Brooks in 1999. It shows similarities to the brachiosaurid ''
Eucamerotus ''Eucamerotus'' (meaning "well-chambered", in reference to the hollows of the vertebrae) was a genus of sauropod dinosaur from the Barremian-age Lower Cretaceous Wessex Formation ( Wealden) of the Isle of Wight, England. History and taxonomy J ...
'' from the
Wessex Formation The Wessex Formation is a fossil-rich English geological formation that dates from the Berriasian to Barremian stages (about 145–125 million years ago) of the Early Cretaceous. It forms part of the Wealden Group and underlies the younger Vect ...
of southern England, as well as to ''
Brachiosaurus ''Brachiosaurus'' () is a genus of sauropod dinosaur that lived in North America during the Late Jurassic, about 154to 150million years ago. It was first described by American paleontologist Elmer S. Riggs in 1903 from fossils found in th ...
'' from the Morrison Formation.


Description

''Cedarosaurus'' was identified as a brachiosaurid sauropod. Prior to its classification as a new genus, brachiosaurid fossil material from the early and middle Cretaceous Period which had been found in North America was ascribed to the genus ''Pleurocoelus'', which is now regarded as a junior synonym of ''
Astrodon ''Astrodon'' (aster: star, odon: tooth) is a genus of large herbivorous sauropod dinosaur, measuring in length, in height and in body mass. It lived in what is now the eastern United States during the Early Cretaceous period, and fossils have ...
''. However, the shape of the animal's vertebrae and forelimb bones were distinct enough to warrant its classification as a new genus. ''Cedarosaurus'' had a more gracile
ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...
and radius than its relative ''
Venenosaurus ''Venenosaurus'' ( ) was a sauropod dinosaur. The name literally means "poison lizard", and it was named so after the Poison Strip Member of the Cedar Mountain Formation in Utah, United States, where the fossils were discovered by a Denver Museum ...
''. The ratio of the radius' least circumference to its length is .31 in ''Cedarosaurus''. Metatarsal II is more gracile in ''Cedarosaurus''. Its middle tail vertebrae's
neural spines The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
are angled anteriorly when the vertebrae are aligned. These vertebrae resemble those of '' Gondwanatitan'', ''
Venenosaurus ''Venenosaurus'' ( ) was a sauropod dinosaur. The name literally means "poison lizard", and it was named so after the Poison Strip Member of the Cedar Mountain Formation in Utah, United States, where the fossils were discovered by a Denver Museum ...
'', and ''
Aeolosaurus ''Aeolosaurus'' (; "Aeolus' lizard") is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. Like most sauropods, it would have been a quadrupedal herbivore with a long neck and tail. ''Aeolos ...
''. The related ''Venenosaurus'' had unusual lateral fossae, which looked like deep depressions in the outside walls of the vertebral centra. Some fossae are divided into two chambers by a ridge inside the depression. In most sauropods the fossae would form pneumatic openings leading to the interior of the centrum, rather than just being a depression. Less well-developed, but similar fossae are known from ''Cedarosaurus'' itself. In 2010, Gregory S. Paul estimated the maximum length up to and mass up to .


Gastroliths

In 2001 Frank Sanders, Kim Manley, and
Kenneth Carpenter Kenneth Carpenter (born September 21, 1949, in Tokyo, Japan) is a paleontologist. He is the former director of the USU Eastern Prehistoric Museum and author or co-author of books on dinosaurs and Mesozoic life. His main research interests ...
published a study on 115
gastrolith A gastrolith, also called a stomach stone or gizzard stone, is a rock held inside a gastrointestinal tract. Gastroliths in some species are retained in the muscular gizzard and used to grind food in animals lacking suitable grinding teeth. In oth ...
s discovered in association with a ''Cedarosaurus'' specimen. The stones were identified as gastroliths on the basis of their tight spatial distribution, partial matrix support, and an edge-on orientation indicative of their being deposited while the carcass still had soft tissue. Their high surface
reflectance The reflectance of the surface of a material is its effectiveness in reflecting radiant energy. It is the fraction of incident electromagnetic power that is reflected at the boundary. Reflectance is a component of the response of the electronic ...
values are consistent with other known dinosaur gastroliths. Nearly all of the ''Cedarosaurus'' gastroliths were found within a .06 m volume of space in the gut region of the skeleton. The total mass of the gastroliths themselves was . Most were less than in volume. The least massive clast was and the most was , with most of them being toward the smaller end of that range. The clasts tended to be close to spherical in shape, although the largest specimens were also the most irregular. The largest gastroliths contributed the most to the total surface area of the set. Some gastroliths were so large and irregularly shaped that they may have been difficult to swallow. The gastroliths were mostly composed of
chert Chert () is a hard, fine-grained sedimentary rock composed of microcrystalline or cryptocrystalline quartz, the mineral form of silicon dioxide (SiO2). Chert is characteristically of biological origin, but may also occur inorganically as a ...
, with some
sandstone Sandstone is a clastic sedimentary rock composed mainly of sand-sized (0.0625 to 2 mm) silicate grains. Sandstones comprise about 20–25% of all sedimentary rocks. Most sandstone is composed of quartz or feldspar (both silicates ...
, siltstone, and
quartzite Quartzite is a hard, non- foliated metamorphic rock which was originally pure quartz sandstone.Essentials of Geology, 3rd Edition, Stephen Marshak, p 182 Sandstone is converted into quartzite through heating and pressure usually related to tec ...
clasts also included. Some of the chert clasts actually contained fossils. Since some of the most irregular gastroliths are also the largest, it is unlikely that they were ingested by accident. ''Cedarosaurus'' may have found irregular clasts to be attractive potential gastroliths or was not selective about shape. The clasts were generally of dull coloration, suggesting that color was not a major factor for the sauropod's decision making. The high surface area to volume ratio of the largest clasts suggests that the gastroliths may have broken down ingested plant material by grinding or crushing it. The sandstone clasts tended to be fragile and some broke in the process of collection. The sandstone gastroliths may have been rendered fragile after deposition by loss of cement caused by the external chemical environment. If the clasts had been that fragile while the animal was alive, they probably rolled and tumbled in the digestive tract. If they were more robust, they could have served as part of a ball-mill system.


Footnotes


References

* * Tidwell, Virginia; Carpenter, Kenneth; Brooks, William (1999)
"New sauropod from the Lower Cretaceous of Utah, USA"
''Oryctos'' 2: 21–37. * Tidwell, V., Carpenter, K. & Meyer, S. (2001). "New Titanosauriform (Sauropoda) from the Poison Strip Member of the Cedar Mountain Formation (Lower Cretaceous), Utah". In: ''Mesozoic Vertebrate Life''. D.H. Tanke & K. Carpenter (eds.). Indiana University Press. pp. 139–165. .


External links

* ''Cedarosaurus'' a
The Paleobiology Database
{{Taxonbar, from=Q134639 Brachiosaurs Early Cretaceous dinosaurs of North America Fossil taxa described in 1999 Taxa named by Kenneth Carpenter Paleontology in Utah