Causus Lichtensteinii
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''Causus lichtensteinii'' is a
viper The Viperidae (vipers) are a family of snakes found in most parts of the world, except for Antarctica, Australia, Hawaii, Madagascar, and various other isolated islands. They are venomous and have long (relative to non-vipers), hinged fangs tha ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to
western Western may refer to: Places *Western, Nebraska, a village in the US *Western, New York, a town in the US *Western Creek, Tasmania, a locality in Australia *Western Junction, Tasmania, a locality in Australia *Western world, countries that id ...
, central, and
eastern Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historica ...
. There are no subspecies that are recognized as being valid. Like all other vipers, it is
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
.


Etymology

The specific name, or epithet, ''lichtensteinii'', honors
German German(s) may refer to: * Germany (of or related to) ** Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ge ...
herpetologist Herpetology (from Greek ἑρπετόν ''herpetón'', meaning "reptile" or "creeping animal") is the branch of zoology concerned with the study of amphibians (including frogs, toads, salamanders, newts, and caecilians ( gymnophiona)) and rep ...
Martin Hinrich Lichtenstein.


Common names

Common names for ''C. lichtensteinii'' include Lichtenstein's night adder, forest night adder,Spawls S, Branch B (1995). ''The Dangerous Snakes of Africa: Natural History, Species Directory, Venoms and Snakebite''. Dubai: Ralph Curtis Publishing. 192 pp. . and olive-green viper.U.S. Navy (1991). ''Poisonous Snakes of the World''. New York: United States Government / Dover Publications Inc. 203 pp. .


Description

Adults of ''C. lichtensteinii'' average in length with a reported maximum of . The head is not very wide, and the snout is blunt. The eye is surrounded by a circumorbital ring of 5–7 scales. There are 6 supralabials and 9 sublabials. The temporals number 2+3 or sometimes 2+2, with the first and second upper temporals being as long together as the first lower one. Loreals: 1+1. Midbody there are 15 rows of weakly keeled
dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down t ...
that have a velvety texture. The
anal scale Anal may refer to: Related to the anus *Related to the anus of animals: ** Anal fin, in fish anatomy ** Anal vein, in insect anatomy ** Anal scale, in reptile anatomy *Related to the human anus: ** Anal sex, a type of sexual activity involv ...
is single. There are 128–152
ventral scales In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...
. The
subcaudals In snakes, the subcaudal scales are the enlarged plates on the underside of the tail.Wright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates (7th printing, 1985). 1105 pp. . These scales may be either single or divided (pair ...
number 18–22 in males and 17–19 in females. The color pattern consists of a greenish or olive ground color overlaid with a series of dark narrow backward pointing chevrons running down the back. This pattern may be vague or developed fully into rhombic markings. The back of the neck has a characteristic white V-shape while the throat is black with yellow bands. Juvenile specimens are generally dark brown in color.


Geographic range

''Causus lichtensteinii'' is found from Guinea and Liberia eastward through
Ghana Ghana (; tw, Gaana, ee, Gana), officially the Republic of Ghana, is a country in West Africa. It abuts the Gulf of Guinea and the Atlantic Ocean to the south, sharing borders with Ivory Coast in the west, Burkina Faso in the north, and To ...
to
Nigeria Nigeria ( ), , ig, Naìjíríyà, yo, Nàìjíríà, pcm, Naijá , ff, Naajeeriya, kcg, Naijeriya officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf o ...
,
Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...
, Equatorial Guinea, the
Central African Republic The Central African Republic (CAR; ; , RCA; , or , ) is a landlocked country in Central Africa. It is bordered by Chad to the north, Sudan to the northeast, South Sudan to the southeast, the DR Congo to the south, the Republic of th ...
, south to northern
Angola , national_anthem = " Angola Avante"() , image_map = , map_caption = , capital = Luanda , religion = , religion_year = 2020 , religion_ref = , coordina ...
, DR Congo and northwestern
Zambia Zambia (), officially the Republic of Zambia, is a landlocked country at the crossroads of Central, Southern and East Africa, although it is typically referred to as being in Southern Africa at its most central point. Its neighbours are t ...
, and east to
Uganda }), is a landlocked country in East Africa. The country is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the south by Tanzania. The sou ...
and western
Kenya ) , national_anthem = " Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi ...
. The type locality is listed as "''Côte- d'Or'' " (
Gold Coast Gold Coast may refer to: Places Africa * Gold Coast (region), in West Africa, which was made up of the following colonies, before being established as the independent nation of Ghana: ** Portuguese Gold Coast (Portuguese, 1482–1642) ** Dutch G ...
, Ghana).Jan (1859).


Habitat

As opposed to other members of its
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
, ''C. lichtensteinii'' is mostly found in pristine rain forests with little light filtering down to the forest floor. It tends to be found near water in swampy areas. In the
Atewa Range The Atewa Range Forest Reserve (also called the Atiwa-Atwaredu ranges) is in the Akyem Abuakwa region of southeastern Ghana, near the town of Kibi, and south-west of the Kwahu Plateau which forms the south-west boundary of Lake Volta. The range ...
Forest Reserve in
Ghana Ghana (; tw, Gaana, ee, Gana), officially the Republic of Ghana, is a country in West Africa. It abuts the Gulf of Guinea and the Atlantic Ocean to the south, sharing borders with Ivory Coast in the west, Burkina Faso in the north, and To ...
it has been found at altitudes of up to .


Behavior

''Causus lichtensteinii'' is diurnal and mostly
terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
. However, it is a good swimmer and has even colonized certain islands in Lake Victoria. When disturbed it puts on a hissing and puffing threat display similar to other members of the genus.Mallow D, Ludwig D, Nilson G (2003). ''True Vipers: Natural History and Toxinology of Old World Vipers''. Malabar, Florida: Krieger Publishing Company. 359 pp. .


Reproduction

''Causus lichtensteinii'' is oviparous.


References


Further reading

* Boulenger GA (1896). ''Catalogue of the Snakes in the British Museum (Natural History). Volume III. Containing the ... Viperidæ''. London: Trustees of the British Museum (Natural History). (Taylor and Francis, printers). xiv + 727 pp. + Plates I-XXV. (''Causus lichtensteinii'', p. 470). * Jan (1859). "''Additions et rectifications aux'' Plan ''et'' Prodrome ''de l' ''Iconographie descriptive des Ophidiens". ''Revue et magasin de zoologie pure et appliquée, 2e serie'' 11: 505-512. (''Aspidelaps lichtensteinii'', new species, p. 511). (in French). {{Taxonbar, from=Q1513923 lichtensteinii Snakes of Africa Reptiles of West Africa Reptiles of Angola Reptiles of Cameroon Reptiles of the Central African Republic Reptiles of the Democratic Republic of the Congo Reptiles of Gabon Reptiles of Kenya Reptiles of Nigeria Reptiles of the Republic of the Congo Reptiles of South Sudan Reptiles of Uganda Reptiles of Zambia Taxa named by Giorgio Jan Reptiles described in 1859