Causus Bilineatus
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''Causus bilineatus'' is a
viper The Viperidae (vipers) are a family of snakes found in most parts of the world, except for Antarctica, Australia, Hawaii, Madagascar, and various other isolated islands. They are venomous and have long (relative to non-vipers), hinged fangs tha ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to south
central Africa Central Africa is a subregion of the African continent comprising various countries according to different definitions. Angola, Burundi, the Central African Republic, Chad, the Democratic Republic of the Congo, the Republic of the Congo ...
. No subspecies are currently recognized. Common names include lined night adder,Mallow D, Ludwig D, Nilson G. 2003. ''True Vipers: Natural History and Toxinology of Old World Vipers''. Krieger Publishing Company. Malabar, Florida. 359 pp. . two-lined night adder, and two-striped night adder.Spawls S, Branch B. 1995. ''The Dangerous Snakes of Africa''. Ralph Curtis Books. Dubai: Oriental Press. 192 pp. . Like all other vipers, it is
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
.


Description

The average total length (body + tail) is with a reported maximum of . The head is slightly distinct from the neck, while the snout is fairly long and tapering. Midbody there are 15–18 rows of
dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down t ...
that are weakly keeled and have a soft and velvety appearance. The
ventral scales In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...
number 122–141 in males and 128–144 in females. There are 18–30
subcaudals In snakes, the subcaudal scales are the enlarged plates on the underside of the tail.Wright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates (7th printing, 1985). 1105 pp. . These scales may be either single or divided (pair ...
. The color pattern consists of an ash to auburn to brown ground color, overlaid with numerous irregular or vaguely rectangular black dorsal patches. These patches lie within two distinct and narrow pale stripes that run the length of the body. The belly color is dark to dark cream.


Geographic range

''Causus bilineatus'' is found in
Angola , national_anthem = " Angola Avante"() , image_map = , map_caption = , capital = Luanda , religion = , religion_year = 2020 , religion_ref = , coordina ...
,
Zambia Zambia (), officially the Republic of Zambia, is a landlocked country at the crossroads of Central, Southern and East Africa, although it is typically referred to as being in Southern Africa at its most central point. Its neighbours are t ...
, southern DR Congo, Rwanda, and western
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands ...
. The type locality is given by Boulenger (1905) as "between
Benguella Benguela (; Umbundu: Luombaka) is a city in western Angola, capital of Benguela Province. Benguela is one of Angola's most populous cities with a population of 555,124 in the city and 561,775 in the municipality, at the 2014 census. History Por ...
and Bihé" (Angola).
Bocage Bocage (, ) is a terrain of mixed woodland and pasture characteristic of parts of Northern France, Southern England, Ireland, the Netherlands and Northern Germany, in regions where pastoral farming is the dominant land use. ''Bocage'' may als ...
(1895) listed "Duque de Bragança, Quissange, Caconda, and Huilla" (Angola) for the localities.


Habitat

Occurs in moist
savanna A savanna or savannah is a mixed woodland- grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
, forest-savanna
environment Environment most often refers to: __NOTOC__ * Natural environment, all living and non-living things occurring naturally * Biophysical environment, the physical and biological factors along with their chemical interactions that affect an organism or ...
s, and swampy
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s. Wild-caught specimens found to have eaten clawed frogs, ''
Xenopus ''Xenopus'' () (Gk., ξενος, ''xenos''=strange, πους, ''pous''=foot, commonly known as the clawed frog) is a genus of highly aquatic frogs native to sub-Saharan Africa. Twenty species are currently described within it. The two best-know ...
'', suggest a more aquatic nature than other species.


Feeding

Preys Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
mostly upon
toad Toad is a common name for certain frogs, especially of the family Bufonidae, that are characterized by dry, leathery skin, short legs, and large bumps covering the parotoid glands. A distinction between frogs and toads is not made in scient ...
s and
frog A frog is any member of a diverse and largely carnivorous group of short-bodied, tailless amphibians composing the order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-frog" ''Triadobatrachus'' is ...
s, particularly the clawed frog, ''Xenopus''.


References


Further reading

* Bocage, J.V.B. du. 1895. ''Herpétologie d'Angola et du Congo''. Ministère de la Marine et des Colonies. (Imprimerie National, printer.) Lisbon. xx + 203 pp. + Plates I-XIX. (''Causus rhombeatus'', part, p. 146.) * Boulenger, G.A. 1905. A List of the Batrachians and Reptiles collected by Dr. W.J. Ansorge in Angola, with Descriptions of new Species. Ann. Mag. Nat. Hist., Seventh Series 16: 105-1156. (''Causus rhombeatus'' var. ''bilineatus'', p. 114.) {{Taxonbar, from=Q233142 bilineatus Snakes of Africa Reptiles of Angola Reptiles of the Democratic Republic of the Congo Vertebrates of Rwanda Reptiles of Tanzania Reptiles of Zambia Reptiles described in 1905 Taxa named by George Albert Boulenger