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The Caspian whipsnake (''Dolichophis caspius'', sometimes also ''Coluber caspius'') also known as the large whipsnake (among various other species in genus ''Dolichophis''/'' Coluber''), is a common species of
whipsnake ''Masticophis'' is a genus of colubrid snakes, commonly referred to as whip snakes or coachwhips, which are endemic to the Americas. Conant R (1975). ''A Field Guide to Reptiles and Amphibians of Eastern and Central North America, Second Edition ...
found in the
Balkans The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throughout the who ...
and parts of
Eastern Europe Eastern Europe is a subregion of the Europe, European continent. As a largely ambiguous term, it has a wide range of geopolitical, geographical, ethnic, cultural, and socio-economic connotations. The vast majority of the region is covered by Russ ...
.


Description

The Caspian whipsnake is perhaps the largest species of snake in Europe. It typically grows to around in length, though a few may exceed a length of . The record sized specimen was approximately Body mass can be commonly from . This species is not venomous and is mostly active during the day. The head is joined to the body by a thick neck. Its pupils are round. Nineteen rows of smooth scales can be found at the middle body, though rarely there may be seventeen. Dorsal scales each have two apical pits at the posterior edge. The centre of the scale is lighter in colour than the edges. The
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal co ...
side is grey-brown and features markings which are distinctive in juveniles and young snakes but fade with age. The ventral side is light yellow or white.


Taxonomy


Synonyms

Since its original description by
Johann Friedrich Gmelin , fields = , workplaces = University of GöttingenUniversity of Tübingen , alma_mater = University of Tübingen , doctoral_advisor = Philipp Friedrich GmelinFerdinand Christoph Oetinger , academic_advisors = , doctora ...
in 1789, ''D. caspius'' has been known by many names. A list of such
synonym A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
s is given below: *''Coluber caspius'' *''Coluber acontistes'' *''Coluber thermalis'' *''Coluber jugularis caspius'' *''Hierophis caspius'' *''Dolichophis caspius'' ''
Nota bene (, or ; plural form ) is a Latin phrase meaning "note well". It is often abbreviated as NB, n.b., or with the ligature and first appeared in English writing . In Modern English, it is used, particularly in legal papers, to draw the atten ...
'': In a synonymy, names without dashes are new
taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
(described as new species). However, names with dashes are just new combinations (moved to a different genus, changed from species to subspecies, etc.).


Distribution

The Caspian whipsnake is found in the
Balkan peninsula The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throughout the who ...
, parts of Eastern Europe and a small portion of Asia Minor. It may be found in
Albania Albania ( ; sq, Shqipëri or ), or , also or . officially the Republic of Albania ( sq, Republika e Shqipërisë), is a country in Southeastern Europe. It is located on the Adriatic and Ionian Seas within the Mediterranean Sea and shares ...
,
Bulgaria Bulgaria (; bg, България, Bǎlgariya), officially the Republic of Bulgaria,, ) is a country in Southeast Europe. It is situated on the eastern flank of the Balkans, and is bordered by Romania to the north, Serbia and North Macedon ...
,
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders with ...
,
North Macedonia North Macedonia, ; sq, Maqedonia e Veriut, (Macedonia before February 2019), officially the Republic of North Macedonia,, is a country in Southeast Europe. It gained independence in 1991 as one of the successor states of Socialist Feder ...
,
Serbia Serbia (, ; Serbian language, Serbian: , , ), officially the Republic of Serbia (Serbian language, Serbian: , , ), is a landlocked country in Southeast Europe, Southeastern and Central Europe, situated at the crossroads of the Pannonian Bas ...
,
Romania Romania ( ; ro, România ) is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern, and Southeast Europe, Southeastern Europe. It borders Bulgaria to the south, Ukraine to the north, Hungary to the west, S ...
,
Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = "Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capit ...
,
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...
,
Moldova Moldova ( , ; ), officially the Republic of Moldova ( ro, Republica Moldova), is a Landlocked country, landlocked country in Eastern Europe. It is bordered by Romania to the west and Ukraine to the north, east, and south. The List of states ...
,
Montenegro ) , image_map = Europe-Montenegro.svg , map_caption = , image_map2 = , capital = Podgorica , coordinates = , largest_city = capital , official_languages = M ...
, southern
Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian inv ...
, southern
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
,
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...
, and southern
Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia a ...
. Results from a survey indicate their habitats may be distributed along low areas near major rivers, such as the
Danube The Danube ( ; ) is a river that was once a long-standing frontier of the Roman Empire and today connects 10 European countries, running through their territories or being a border. Originating in Germany, the Danube flows southeast for , pa ...
and the
Olt River The Olt (Romanian and Hungarian; german: Alt; la, Aluta or ', tr, Oltu, grc, Ἄλυτος ''Alytos'') is a river in Romania. It is long, and its basin area is . It is the longest river flowing exclusively through Romania. Its average disch ...
. It was previously assumed extinct in
Moldavia Moldavia ( ro, Moldova, or , literally "The Country of Moldavia"; in Romanian Cyrillic: or ; chu, Землѧ Молдавскаѧ; el, Ἡγεμονία τῆς Μολδαβίας) is a historical region and former principality in Centr ...
(eastern Romania, southern Ukraine, and western Moldova), where it was only known from two sites, and not observed since 1937. Three specimens were collected in May 2007 in
Galați County Galați () is a county ( județ) of Romania, in Moldavia region, with the capital city at Galați. History Historically Galați is part of Moldavia. In 1858, it was represented by Alexandru Ioan Cuza at the ''ad hoc Divan'' at Iași, in the wa ...
, somewhat dispelling this belief. Though it is known to be common in the
Dobrudja Dobruja or Dobrudja (; bg, Добруджа, Dobrudzha or ''Dobrudža''; ro, Dobrogea, or ; tr, Dobruca) is a historical region in the Balkans that has been divided since the 19th century between the territories of Bulgaria and Romania. I ...
area, knowledge of its distribution in other areas is poor. It is believed to be very rare in these parts, and may also be highly threatened. National legislation has declared ''D. caspius'' to be "a species of community interest" and thus must be strictly protected. In Hungary, it was formerly thought only to exist in two areas: one near the capital of
Budapest Budapest (, ; ) is the capital and most populous city of Hungary. It is the ninth-largest city in the European Union by population within city limits and the second-largest city on the Danube river; the city has an estimated population ...
, and another near Szársomlyó. However, a recent survey of the region has yielded several previously unknown whipsnake habitats along the River Danube, including sites near
Dunaújváros Dunaújváros (; also known by other alternative names) is an industrial city in Fejér County, Central Hungary. It is a city with county rights. Situated 70 kilometres (43 miles) south of Budapest on the Danube, the city is best known for its ...
and
Dunaföldvár Dunaföldvár is a town in Tolna County, Hungary. Residents are Hungarians, with minority of Serbs. A Bronze Age gold hoard of jewellery was found between Paks and Dunaföldvár on the banks of the Danube in the nineteenth century. The treasure ...
. As in Romania, ''D. caspius'' is a protected species in Hungary.


Habitat

Recently observed specimens of ''D. caspius'' were found in grassy, open areas with sparse
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
bery. They have also been found in open, bushy areas, specifically, between the Danube Meadow and
Romanian Plain The Romanian Plain ( ro, Câmpia Română) is located in southern Romania and the easternmost tip of Serbia, where it is known as the Wallachian Plain ( sr, Vlaška nizija/Влашка низија). Part of the historical region of Wallachia, it ...
, near Căscioarele. Vine covered areas were found near this habitat as well. At Drăgăneşti Olt, the species was found on a hilly slope. Its habitat also includes rocky riverbanks and swampy areas, as well as gardens and cemeteries.


Diet

The Caspian whipsnake is a generalist predator and feeds on various vertebrates. Its diet may include small mammals, lizards, birds, and other snakes. Though not venomous, the Caspian whipsnake is considered "ferocious" and bites quickly and without warning. Due to its ability to consume rodents which may harm crops, it is valued as natural
pest control Pest control is the regulation or management of a species defined as a pest; any animal, plant or fungus that impacts adversely on human activities or environment. The human response depends on the importance of the damage done and will range ...
.


Conservation

Data on the conservation status of ''D. caspius'' in many regions are sparse. Whilst it is known to be common in the
Dobrudja Dobruja or Dobrudja (; bg, Добруджа, Dobrudzha or ''Dobrudža''; ro, Dobrogea, or ; tr, Dobruca) is a historical region in the Balkans that has been divided since the 19th century between the territories of Bulgaria and Romania. I ...
area, it may be rare and threatened in other areas. In Drăgăneşti Olt, Romania, road-killed snakes are a "common sight" to locals. Traffic-related deaths may be a cause of their decline in Romania. One of the recently observed habitats, the
Romanian Plain The Romanian Plain ( ro, Câmpia Română) is located in southern Romania and the easternmost tip of Serbia, where it is known as the Wallachian Plain ( sr, Vlaška nizija/Влашка низија). Part of the historical region of Wallachia, it ...
, is known to be heavily affected by agriculture.
Habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
to agriculture may also be a cause of the species' decline in Romania.


References

{{Taxonbar, from=Q41694 Dolichophis Reptiles described in 1789 Taxa named by Johann Friedrich Gmelin Snakes of Jordan Reptiles of Europe Reptiles of Russia