Carissa Spinarum
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''Carissa spinarum'', the conkerberry or bush plum, is a large
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
of the dogbane
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
(
Apocynaceae Apocynaceae (from ''Apocynum'', Greek for "dog-away") is a family of flowering plants that includes trees, shrubs, herbs, stem succulents, and vines, commonly known as the dogbane family, because some taxa were used as dog poison Members of the ...
), widely distributed in tropical regions of Africa, Southern Asia, Australia, and various islands of the Indian Ocean. It is most well known in Australia, where it is also called ''currant bush'' or, more ambiguously, ''native currant'' or even ''black currant''. It is, however, neither closely related to
plum A plum is a fruit of some species in ''Prunus'' subg. ''Prunus'.'' Dried plums are called prunes. History Plums may have been one of the first fruits domesticated by humans. Three of the most abundantly cultivated species are not found i ...
s (''Prunus'') nor to true currants (''Ribes''), which belong to entirely different lineages of
eudicot The eudicots, Eudicotidae, or eudicotyledons are a clade of flowering plants mainly characterized by having two seed leaves upon germination. The term derives from Dicotyledons. Traditionally they were called tricolpates or non-magnoliid dicot ...
s. In
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
, it is also called wild karanda /wild karavanda, referring to the related karanda (''C. carandas''). ''Carissa spinarum'' is often discussed under its many obsolete
synonym A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
s (see below). It grows as a multi-stemmed shrub, 0.5 to 3 metres in height. The leaves are glossy green, opposite, narrow ovate to lanceolate and 1–5 cm in length. The branches bear thorns of 1–3 cm length. White, star-shaped flowers ~1 cm across are followed by ovate green berries, 1–2 cm in length, which turn black or dark purple when ripe.


Ecology

''Carissa spinarum'' is most often found in
semiarid A semi-arid climate, semi-desert climate, or steppe climate is a dry climate sub-type. It is located on regions that receive precipitation below potential evapotranspiration, but not as low as a desert climate. There are different kinds of semi-ar ...
coastal regions on fine-textured soils such as
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolin, Al2 Si2 O5( OH)4). Clays develop plasticity when wet, due to a molecular film of water surrounding the clay par ...
s and clay-
loam Loam (in geology and soil science) is soil composed mostly of sand (particle size > ), silt (particle size > ), and a smaller amount of clay (particle size < ). By weight, its mineral composition is about 40–40–20% concentration of sand–sil ...
s; in more
arid A region is arid when it severely lacks available water, to the extent of hindering or preventing the growth and development of plant and animal life. Regions with arid climates tend to lack vegetation and are called xeric or desertic. Most ar ...
regions the plant tends to be confined to areas of higher moisture such as at the base of hills or floodout areas. But it has a high ecological tolerance and can live in a wide range of
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s. In Australia for example, it is often found in association with ''
Eucalyptus brownii ''Eucalyptus'' () is a genus of over seven hundred species of flowering trees, shrubs or mallees in the myrtle family, Myrtaceae. Along with several other genera in the tribe Eucalypteae, including ''Corymbia'', they are commonly known as euc ...
'',
poplar box ''Eucalyptus populnea'', commonly known as poplar box, bimble box or bimbil box, is a species of small to medium-sized tree that is endemic to eastern Australia. It has rough, fibrous or flaky bark on the trunk and branches, egg-shaped, elliptic ...
(''E. populnea''), gidgee (''Acacia cambagei'') or
brigalow ''Acacia harpophylla'', commonly known as brigalow, brigalow spearwood or orkor, is an endemic tree of Australia. The Aboriginal Australian group the Gamilaraay peoples know the tree as Barranbaa or Burrii. It is found in central and coasta ...
(''A. harpophylla''), in coastal rainforest, gallery forest and vine thickets in regions receiving in excess of 900 mm annual rainfall, as well as softwood scrubs and open
eucalypt Eucalypt is a descriptive name for woody plants with capsule fruiting bodies belonging to seven closely related genera (of the tribe Eucalypteae) found across Australasia: ''Eucalyptus'', '' Corymbia'', '' Angophora'', ''Stockwellia'', ''Allosyn ...
savannas receiving less than 700 mm annual rainfall. Conkerberries are edible, but only when fully ripe; they have a sweet flavour, but the milky sap of this plant – and its unripe fruit – is poisonous, as typical for the Apocynaceae. They are a popular
bush tucker Bush tucker, also called bush food, is any food native to Australia and used as sustenance by Indigenous Australians, the Aboriginal and Torres Strait Islander peoples, but it can also describe any native flora or fauna used for culinary or ...
food for
Australian Aborigines Aboriginal Australians are the various Indigenous peoples of the Australian mainland and many of its islands, such as Tasmania, Fraser Island, Hinchinbrook Island, the Tiwi Islands, and Groote Eylandt, but excluding the Torres Strait Islands ...
in Central Australia. The fruit is known as ''merne arrankweye'' in the
Arrernte language Arrernte or Aranda (; ) or sometimes referred to as Upper Arrernte (Upper Aranda), is a dialect cluster in the Arandic language group spoken in parts of the Northern Territory, Australia, by the Arrernte people. Other spelling variations are A ...
, ''anwekety'' in
Anmatyerr The Anmatyerr, also spelt Anmatyerre, Anmatjera, Anmatjirra, Amatjere and other variations) are an Aboriginal Australian people of the Northern Territory, who speak one of the Upper Arrernte languages. Language Anmatyerr is divided into Easte ...
and ''nganango'' in
Pintupi The Pintupi are an Australian Aboriginal group who are part of the Western Desert cultural group and whose traditional land is in the area west of Lake Macdonald and Lake Mackay in Western Australia. These people moved (or were moved) into the ...
. The fruits are also a popular food for the
Australian bustard The Australian bustard (''Ardeotis australis'') is a large ground dwelling bird which is common in grassland, woodland and open agricultural country across northern Australia and southern New Guinea. It stands at about high, and its wingspan is ...
,
emu The emu () (''Dromaius novaehollandiae'') is the second-tallest living bird after its ratite relative the ostrich. It is endemic to Australia where it is the largest native bird and the only extant member of the genus ''Dromaius''. The emu' ...
and many other birds in its range. Its leaves provide food for butterflies (e.g.
Australian crow The Torresian crow (''Corvus orru''), also called the Australian crow or Papuan crow, is a passerine bird in the crow family native to the north and west of Australia and nearby islands in Indonesia and Papua New Guinea. The species has a black p ...
, ''Euploea core'') and moths (e.g. some
hawkmoth The Sphingidae are a family of moths ( Lepidoptera) called sphinx moths, also colloquially known as hawk moths, with many of their caterpillars known as “hornworms”; it includes about 1,450 species. It is best represented in the tropics, b ...
s) ''Carissa spinarum'' is frequently a weed in grazing land in northern Australia, choking out grasses, reducing the ability of livestock to feed, interfering with stock handling and providing a refuge for vermin. The plant is capable of reproducing rapidly by
layering Layering has evolved as a common means of vegetative propagation of numerous species in natural environments. Layering is also utilized by horticulturists to propagate desirable plants. Natural layering typically occurs when a branch touches ...
and is difficult to control mechanically and expensive to manage with
herbicides Herbicides (, ), also commonly known as weedkillers, are substances used to control undesired plants, also known as weeds.EPA. February 201Pesticides Industry. Sales and Usage 2006 and 2007: Market Estimates. Summary in press releasMain page fo ...
. On the other hand, it has been used in attempts to restore small-bird habitat in disturbed dry rainforest in Queensland, Australia. Parts of the plant are used medicinally for joint and muscle pain by the
Maasai people The Maasai (; sw, Wamasai) are a Nilotic ethnic group inhabiting northern, central and southern Kenya and northern Tanzania. They are among the best-known local populations internationally due to their residence near the many game parks of t ...
of
Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...
.


Synonyms

Well known for its fruit to locals and quite variable across its wide range and diverse habitat types, the conkerberry has been described time and again by
botanist Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...
s under a number of names. Robert Brown alone described it no less than four times under different names, and R.H. Beddome not only described it twice as a "new
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
" of ''
Carissa ''Carissa'' is a genus of shrubs or small trees native to tropical and subtropical regions of Africa, Australia and Asia. Until recently about 100 species were listed, but most of them have been relegated to the status of synonyms or assigned ...
'', but believed two other growth-forms of it to be mere
varieties Variety may refer to: Arts and entertainment Entertainment formats * Variety (radio) * Variety show, in theater and television Films * ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont * ''Variety'' (1935 film), ...
of Karonda (''C. carandas''). The 1889 book 'The Useful Native Plants of Australia records the plant under the name Carissa Ovata and says "This little bush produces a very pleasant fruit, which is both agreeable and wholesome. It is like a sloe, egg-shaped, and about half-an-inch long. It exudes a viscid milky juice and contains a few woody seeds. "I can testify that the fruit is both agreeable and wholesome, and I never knew an instance of any evil consequences, even when they were partaken of most abundantly." (Tenison-Woods, Vol. vii., 571., Proc. Lirin. Soc. N.S.W.)". But in fact, ''C. spinarum'' was already named by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in his 1771 ''
Mantissa Plantarum Mantissa () may refer to: * Mantissa (logarithm), the fractional part of the common (base-10) logarithm * Mantissa (floating point number), the significant digits of a floating-point number or a number in scientific notation, also called the ''sig ...
'', and thus all subsequent names are treated as
junior synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linna ...
s. In Myanmar, it is known as . The following list gives the names under which the conkerberry has been placed in ''Carissa''. Apart from that, it has also been assigned, under various names, to ''Antura'' and ''Arduina'' (both now synonymized with ''Carissa''), as well as ''
Azima ''Azima'' is a genus of plants in the family Salvadoraceae. Species include *'' Azima angustifolia'' *'' Azima sarmentosa'' *''Azima tetracantha ''Azima tetracantha'' ( syn. ''Monetia barlerioides'' L'Her.) is an ornamental plant in the Sal ...
'', '' Cabucala'', '' Chapelieria'', ''
Damnacanthus ''Damnacanthus'' is a genus of flowering plants in the family Rubiaceae. The genus is found from Assam to temperate eastern Asia. Species * '' Damnacanthus angustifolius'' Hayata * '' Damnacanthus biflorus'' (Rehder) Masam. * '' Damnacanthus gig ...
'', ''
Strychnos ''Strychnos'' is a genus of flowering plants, belonging to the family Loganiaceae (sometimes Strychnaceae). The genus includes about 100 accepted species of trees and lianas, and more than 200 that are as yet unresolved. The genus is widely dis ...
'', ''Carandas'', and ''Jasminonerium''.Tropicos.org
009 009 may refer to: * OO9, gauge model railways * O09, FAA identifier for Round Valley Airport * 0O9, FAA identifier for Ward Field, see List of airports in California * British secret agent 009, see 00 Agent * BA 009, see British Airways Flight 9 * ...
/ref>
* ''Carissa abyssinica'' R.Br. * ''Carissa africana'' A.DC. * ''Carissa axillaris'' Roxb. * ''Carissa brownii'' F.Muell. * ''Carissa campenonii'' (Drake) Palacky * ''Carissa candolleana'' Jaub. & Spach * ''Carissa carandas'' var. ''congesta'' (Wight) Bedd. * ''Carissa carandas'' var. ''paucinervia'' (A.DC.) Bedd. * ''Carissa cochinchinensis'' Pierre ex Pit. * ''Carissa comorensis'' (Pichon) Markgr. * ''Carissa congesta'' Wight * ''Carissa coriacea'' Wall. * ''Carissa cornifolia'' Jaub. & Spach * ''Carissa dalzellii'' Bedd. * ''Carissa densiflora'' Baker * ''Carissa diffusa'' Roxb. * ''Carissa dulcis'' Schumach. & Thonn. * ''Carissa edulis'' (Forssk.) Vahl :''C. edulis'' var. ''septentrionalis'' is probably a valid species, '' Carissa septentrionalis'' * ''Carissa hirsuta'' Roth * ''Carissa horrida'' Pichon * ''Carissa inermis'' Vahl * ''Carissa lanceolata'' R.Br. * ''Carissa laxiflora'' Benth. * ''Carissa macrophylla'' Wall. * ''Carissa madagascariensis'' Thouars ex Poir. * ''Carissa obovata'' Markgr. * ''Carissa oleoides'' Markgr. * ''Carissa ovata'' R.Br. * ''Carissa paucinervia'' A.DC. * ''Carissa pilosa'' Schinz * ''Carissa pubescens'' A.DC. * ''Carissa revoluta'' Scott-Elliot * ''Carissa richardiana'' Jaub. & Spach * ''Carissa scabra'' R.Br. * ''Carissa sechellensis'' Baker * ''Carissa suavissima'' Bedd. ex Hook.f. * ''Carissa tomentosa'' A.Rich. * ''Carissa villosa'' Roxb. * ''Carissa xylopicron'' Thouars * ''Carissa yunnanensis'' Tsiang & P.T.Li


Footnotes


References

*
009 009 may refer to: * OO9, gauge model railways * O09, FAA identifier for Round Valley Airport * 0O9, FAA identifier for Ward Field, see List of airports in California * British secret agent 009, see 00 Agent * BA 009, see British Airways Flight 9 * ...
br>Synonyms of ''Carissa spinarum'' L.
Retrieved 2009-NOV-26.


External links


Alice Springs Town Council


* {{Taxonbar, from=Q2938847 spinarum Bushfood Australian Aboriginal bushcraft Fruits originating in Africa Flora of Australia Flora of tropical Asia Plants described in 1771 Taxa named by Carl Linnaeus