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The cardueline finches are a
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
, Carduelinae, one of three subfamilies of the finch family
Fringillidae The true finches are small to medium-sized passerine birds in the family Fringillidae. Finches have stout conical bills adapted for eating seeds and nuts and often have colourful plumage. They occupy a great range of habitats where they are usua ...
, the others being the
Fringillinae The genus ''Fringilla'' is a small group of finches from the Old World, which are the only species in the subfamily Fringillinae. The genus name ''Fringilla'' is Latin for "finch". Taxonomy The genus ''Fringilla'' was introduced in 1758 by the ...
and the
Euphoniinae Euphoniinae is a subfamily of finches endemic to the Neotropics. It contains two genera, ''Euphonia'' and '' Chlorophonia''. The two genera were at one time included in the tanager family Thraupidae. In a large phylogenetic study of the finch ...
. The
Hawaiian honeycreeper Hawaiian honeycreepers are a group of small, passerine birds endemic to Hawaii. They are closely related to the rosefinches in the genus ''Carpodacus'', but many species have evolved features unlike those present in any other finch. Their great ...
s are now included in this subfamily. Except for the Hawaiian honeycreepers which underwent
adaptive radiation In evolutionary biology, adaptive radiation is a process in which organisms diversify rapidly from an ancestral species into a multitude of new forms, particularly when a change in the environment makes new resources available, alters biotic int ...
in Hawaii and have evolved a broad range of diets, cardueline finches are specialised seed eaters, and unlike most passerine birds, they feed their young mostly on seeds, which are regurgitated.Newton, 1973, p. 31 Besides this, they differ from the other finches in some minor details of their skull. They are adept at opening seeds and clinging to stems, unlike other
granivorous Seed predation, often referred to as granivory, is a type of plant-animal interaction in which granivores (seed predators) feed on the seeds of plants as a main or exclusive food source,Hulme, P.E. and Benkman, C.W. (2002) "Granivory", pp. 132 ...
birds, such as sparrows and buntings, which feed mostly on fallen seeds. Some members of this subfamily are further specialised to feed on a particular type of seed, such as cones in the case of
crossbill The crossbill is a genus, ''Loxia'', of birds in the finch family (Fringillidae), with six species. These birds are characterised by the mandibles with crossed tips, which gives the group its English name. Adult males tend to be red or orange in ...
s. Carduelines forage in flocks throughout the year, rather than keeping territories, and males defend their females rather than a territory or nest. The name Carduelina for the subfamily was introduced by the Irish zoologist
Nicholas Aylward Vigors Nicholas Aylward Vigors (1785 – 26 October 1840) was an Ireland, Irish zoologist and politician. He popularized the classification of birds on the basis of the quinarian system. Early life Vigors was born at Old Leighlin, County Carlow on 17 ...
in 1825. Carduelinae is derived from the Latin name ''carduelis'' and the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Carduelis carduelis'' for a goldfinch, one of the species in the subfamily.


List of genera

The Carduelinae subfamily contains 186 species divided into 49 genera. Of the 186 species, 15 are now extinct; these are the
Bonin grosbeak The Bonin grosbeak or Bonin Islands grosbeak (''Carpodacus ferreorostris'') is an extinct finch. It is one of the diverse bird taxa that are vernacularly called "grosbeaks", but it is not closely related to the grosbeaks ''sensu stricto''. Many ...
and 14 Hawaiian honeycreepers. * ''
Mycerobas The ''Mycerobas'' grosbeaks are a genus of finch in the family Fringillidae The true finches are small to medium-sized passerine birds in the family Fringillidae. Finches have stout conical bills adapted for eating seeds and nuts and often h ...
'' – contains four Asian
grosbeak Grosbeak is a form taxon containing various species of seed-eating passerine birds with large beaks. Although they all belong to the superfamily Passeroidea, these birds are not part of a natural group but rather a polyphyletic assemblage of di ...
s * ''
Hesperiphona ''Hesperiphona'' is a genus in the finch family Fringillidae. The genus was introduced in 1850 by the French naturalist Charles Lucien Bonaparte with the evening grosbeak as the type species. The name combines the Ancient Greek ''hesperos'' meani ...
'' – contains two American grosbeaks, the
evening grosbeak The evening grosbeak (''Hesperiphona vespertina'') is a passerine bird in the finch family Fringillidae found in North America. Taxonomy The IOC checklist and the '' Handbook of the Birds of the World'' place the evening grosbeak and the clos ...
and the
hooded grosbeak The hooded grosbeak (''Hesperiphona abeillei'') is a passerine bird in the finch family found in the highlands of Central America, principally in Mexico and Guatemala. This is a medium large grosbeak with a large bill. The male has a black head ...
* ''
Coccothraustes The hawfinch (''Coccothraustes coccothraustes'') is a passerine bird in the finch family Fringillidae. It is the only species placed in the genus ''Coccothraustes''. Its closest living relatives are the Chinese grosbeak (''Eophona migratoria'') ...
'' – contains a single species, the
hawfinch The hawfinch (''Coccothraustes coccothraustes'') is a passerine bird in the finch family Fringillidae. It is the only species placed in the genus ''Coccothraustes''. Its closest living relatives are the Chinese grosbeak (''Eophona migratoria'') ...
* ''
Eophona The Oriental grosbeaks (''Eophona'') are a genus of finches containing two species: The genus was introduced in 1851 by the English ornithologist and bird artist John Gould John Gould (; 14 September 1804 – 3 February 1881) was an Engl ...
'' – contains the two oriental grosbeaks, the
Chinese Chinese can refer to: * Something related to China * Chinese people, people of Chinese nationality, citizenship, and/or ethnicity **''Zhonghua minzu'', the supra-ethnic concept of the Chinese nation ** List of ethnic groups in China, people of ...
and the
Japanese grosbeak The Japanese grosbeak (''Eophona personata'') or Ikaru is a finch native to the East Palearctic. It is also sometimes referred to as the Japanese or masked hawfinch due to superficial similarities to the well-known Hawfinch, Eurasian species. ...
* ''
Pinicola The pine grosbeak (''Pinicola enucleator'') is a large member of the true finch family, Fringillidae. It is the only species in the genus ''Pinicola''. It is found in coniferous woods across Alaska, the western mountains of the United States, Can ...
'' – contains a single species, the
pine grosbeak The pine grosbeak (''Pinicola enucleator'') is a large member of the true finch family, Fringillidae. It is the only species in the genus ''Pinicola''. It is found in coniferous woods across Alaska, the western mountains of the United States, Can ...
* ''
Pyrrhula Pyrrhula is a small genus of passerine birds, commonly called bullfinches, belonging to the finch family (Fringillidae). The genus has a Palearctic distribution; almost all species occur in Asia, with two species exclusively in the Himalayas and ...
'' – contains the eight bullfinch species * ''
Rhodopechys Rhodopechys is a genus of finches containing the following two species: The desert finch, ''Carduelis obsoleta'' (formerly ''Rhodopechys obsoleta''), has turned out to belong to the (sub)genus ''Chloris'' in the genus ''Carduelis'' as indicate ...
'' – contains two species, the
Asian crimson-winged finch The Asian crimson-winged finch (''Rhodopechys sanguineus'') is a pale-colored thickset finch with a heavy, dull yellowish bill. It is found from Turkey to NE Pakistan. The African crimson-winged finch was formerly considered conspecific and toge ...
and the
African crimson-winged finch The African crimson-winged finch (''Rhodopechys alienus'') is a pale-colored thickset finch with a heavy, dull yellowish bill. It is found in the Atlas Mountains of Morocco and Algeria. It was formerly considered a subspecies of the Asian crims ...
* '' Bucanetes'' – contains the
trumpeter The trumpet is a brass instrument commonly used in classical and jazz ensembles. The trumpet group ranges from the piccolo trumpet—with the highest register in the brass family—to the bass trumpet, pitched one octave below the standard B ...
and the
Mongolian finch The Mongolian finch (''Bucanetes mongolicus''), also known as the Mongolian trumpeter finch, is a small passerine bird in the finch family Fringillidae. Description The Mongolian finch is a small, long-winged bird. It has a large head and short, ...
* ''
Agraphospiza Blanford's rosefinch (''Agraphospiza rubescens'') or the crimson rosefinch, is a species of finch in the family Fringillidae. It is found in Bhutan, China, India, and Nepal. Its natural habitat is boreal forest. Blanford's rosefinch was formerly ...
'' – contains a single species,
Blanford's rosefinch Blanford's rosefinch (''Agraphospiza rubescens'') or the crimson rosefinch, is a species of finch in the family Fringillidae. It is found in Bhutan, China, India, and Nepal. Its natural habitat is boreal forest. Blanford's rosefinch was formerly ...
* '' Callacanthis'' – contains a single species, the
spectacled finch The spectacled finch (''Callacanthis burtoni'') is a species of finch in the family Fringillidae. It is found in temperate northern regions of the Indian subcontinent, ranging across Afghanistan, India, Nepal, and Pakistan. Its natural habitat is ...
* '' Pyrrhoplectes'' – contains a single species, the
golden-naped finch The golden-naped finch (''Pyrrhoplectes epauletta'') is a species of finch in the family Fringillidae. It is in monotypic genus ''Pyrrhoplectes''. It is found in Bhutan, China, India, Myanmar, and Nepal. Its natural habitat is temperate forest. ...
* '' Procarduelis'' – contains a single species, the
dark-breasted rosefinch The dark-breasted rosefinch (''Procarduelis nipalensis'') is a species of true finch in the monotypic genus ''Procarduelis''. It is found in Bhutan, China, India, Laos, Myanmar, Nepal, Pakistan, Thailand, and Vietnam. Its natural habitats are bor ...
* '' Leucosticte'' – contains six species of mountain and rosy finches * ''
Carpodacus The rosefinches are a genus, ''Carpodacus'', of passerine birds in the finch family Fringillidae. Most are called "rosefinches" and as the word implies, have various shades of red in their plumage. The common rosefinch is frequently called the ...
'' – contains the 28
Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
rosefinch species *
Hawaiian honeycreeper Hawaiian honeycreepers are a group of small, passerine birds endemic to Hawaii. They are closely related to the rosefinches in the genus ''Carpodacus'', but many species have evolved features unlike those present in any other finch. Their great ...
group (formerly a separate family, Drepanididae) ** '' Melamprosops'' – contains a single extinct species, the poo-uli ** ''
Paroreomyza ''Paroreomyza'' is a genus of Hawaiian honeycreeper in the subfamily Carduelinae of the family Fringillidae. These birds are endemic to Hawaii. Taxonomy ''Paroreomyza'', along with '' Oreomystis'' (although their alliance is disputed), is the ...
'' – contains three species, the
Oahu alauahio Oahu () ( Hawaiian: ''Oʻahu'' ()), also known as "The Gathering Place", is the third-largest of the Hawaiian Islands. It is home to roughly one million people—over two-thirds of the population of the U.S. state of Hawaii. The island of O ...
, the
Maui alauahio The island of Maui (; Hawaiian: ) is the second-largest of the islands of the state of Hawaii at 727.2 square miles (1,883 km2) and is the 17th largest island in the United States. Maui is the largest of Maui County's four islands, whic ...
and the extinct kakawahie ** '' Oreomystis'' – contains a single species, the akikiki ** ''
Telespiza ''Telespiza'' is a genus of Hawaiian honeycreeper. All species in it are or were endemic to the Hawaiian Islands. Species *†''Telespiza persecutrix'' James & Olson, 1991 - Kauai finch (prehistoric) *†''Telespiza ypsilon'' James & Olson, ...
'' – contains two species, the
Laysan finch The Laysan finch (''Telespiza cantans'') is a species of Hawaiian honeycreeper, that is endemic to the Northwestern Hawaiian Islands. It is one of four remaining finch-billed Hawaiian honeycreepers and is closely related to the smaller Nihoa fin ...
and the
Nihoa finch The Nihoa finch (''Telespiza ultima'') is one of the two endemic bird species of the tiny Hawaiian island Nihoa, the other being the Nihoa millerbird. When it was classified in 1917, scientists thought that it would be the last endemic species nam ...
** ''
Loxioides ''Loxioides'' is a genus of Hawaiian honeycreeper, in the subfamily Carduelinae. The birds are endemic to Hawaii. Species It contains the following species: * '' Loxioides bailleui'' Oustalet, 1877 - palila * '' Loxioides kikuichi'' James & ...
'' – contains a single species, the
palila The palila (''Loxioides bailleui'') is a critically endangered finch-billed species of Hawaiian honeycreeper. It has a golden-yellow head and breast, with a light belly, gray back, and greenish wings and tail. The bird has a close ecological rel ...
** ''
Rhodacanthis ''Rhodacanthis'' is an extinct genus of Hawaiian honeycreepers commonly known as koa finches. All four species were endemic to the Hawaiian Islands. Members of this genus were granivores, with bills adapted to the seeds and pods of legumes. The ...
'' – contains two extinct species, the
lesser Lesser, from Eliezer (, "Help/Court of my God"), is a surname. Notable people with the surname include: * Adolf Lesser (1851–1926), German physician * Aleksander Lesser (1814–1884), Polish painter and art critic * Anton Lesser (born 1952), Bri ...
and the
greater koa finch The greater koa finch (''Rhodacanthis palmeri'') was a species of finch in the family Fringillidae. It was found only in the Hawaiian Islands. It has been extinct since the late 19th century. Description The bird was about 7–8 in (18–20 ...
** ''
Chloridops ''Chloridops'' is an extinct genus of Hawaiian honeycreeper in the subfamily Carduelinae of the family Fringillidae. Distribution The birds were endemic to Hawaii. It comprised three species: two on the Big Island of Hawaii; and one that inhabi ...
'' – contains a single extinct species, the
Kona grosbeak The Kona grosbeak (''Chloridops kona'') is an extinct species of Hawaiian honeycreeper. The Kona grosbeak was endemic to naio (''Myoporum sandwicense'') forests on ʻaʻā lava flows at elevations of near the Kona District on the island of Hawa ...
** '' Psittirostra'' – contains a single possibly extinct species, the ou ** '' Dysmorodrepanis'' – contains a single extinct species, the
Lanai hookbill Lanai ( haw, Lānai, , , also ,) is the sixth-largest of the Hawaiian Islands and the smallest publicly accessible inhabited island in the chain. It is colloquially known as the Pineapple Island because of its past as an island-wide pineapple pl ...
** ''
Drepanis ''Drepanis'' is a genus of Hawaiian honeycreeper in the subfamily Carduelinae of the family Fringillidae. The birds are endemic to Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific ...
'' – contains two extinct species, the
Hawaii mamo The Hawaii mamo (''Drepanis pacifica'') is an extinct species of Hawaiian honeycreeper. It was endemic to Hawaii. It became extinct due to habitat loss, mosquitoes, introduced predators such as mongoose, and overcollecting. Description This bi ...
and the
black mamo The black mamo (''Drepanis funerea''), also known as the hoa, is an extinction, extinct species of bird once Endemism, endemic to the island of Molokai; there is also subfossil evidence of it having lived on Maui. Description It measured fr ...
, and the extant iiwi ** ''
Ciridops ''Ciridops'' is an extinct genus of Hawaiian honeycreeper species that occurred in prehistoric and historic times on the Hawaiian islands of Hawaii, Molokai, Kauai and Oahu. This genus was created in 1892 by Alfred Newton in an article published ...
'' – contains a single extinct species, the Ula-ai-hawane ** '' Palmeria'' – contains a single species, the akohekohe ** ''
Himatione ''Himatione'' is a genus of Hawaiian honeycreepers in the subfamily Carduelinae of the family Fringillidae. Species It contains the following species: * Laysan honeycreeper (''Himatione fraithii'') (extinct) * ʻApapane The apapane (''Himatio ...
'' – contains two species, the apapane and the extinct
Laysan honeycreeper The Laysan honeycreeper or Laysan apapane (''Himatione fraithii'') is an extinct bird species that was endemic to the island of Laysan in the Northwestern Hawaiian Islands. Taxonomy The species was described by the British ornithologist Walter ...
** '' Viridonia'' – contains a single extinct species, the greater amakihi ** ''
Akialoa ''Akialoa'' is an extinct genus of Hawaiian honeycreeper in the subfamily Carduelinae of the family Fringillidae. The birds were endemic to Hawaii. Species It contains the following species: * Lesser ʻakialoa or Hawai'i ʻakialoa, ''Akialoa ...
'' – contains six extinct species ** ''
Hemignathus ''Hemignathus'' is a Hawaiian honeycreeper genus in the subfamily Carduelinae of the finch family, Fringillidae. These birds are endemic to the Hawaiian Islands. Extinctions Many of its species became extinct during the 19th and 20th centurie ...
'' – contains five species, only one of which is extant, with two being possibly extinct ** '' Pseudonestor'' – contains a single species, the
Maui parrotbill The Maui parrotbill or kiwikiu (''Pseudonestor xanthophrys'') is a species of Hawaiian honeycreeper endemic to the island of Maui in Hawaii. It can only be found in of mesic and wet forests at on the windward slopes of Haleakalā. This spec ...
** '' Magumma'' – contains a single species, the anianiau ** ''
Loxops ''Loxops'' is a Hawaiian honeycreeper genus in the finch family (biology), family, Fringillidae. Most of them are commonly known as akepa. Taxonomy There are 5 species in this genus, two of which are recently extinct or possibly extinct: * ‘A ...
'' – contains five species, of which one is extinct and one possibly extinct ** ''
Chlorodrepanis ''Chlorodrepanis'' is a genus of Hawaiian honeycreeper in the subfamily Carduelinae of the family Fringillidae. The birds are endemic to Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the P ...
'' – contains three species, the
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
,
Oahu Oahu () (Hawaiian language, Hawaiian: ''Oʻahu'' ()), also known as "The Gathering place#Island of Oʻahu as The Gathering Place, Gathering Place", is the third-largest of the Hawaiian Islands. It is home to roughly one million people—over t ...
and Kauai amakihi * ''
Haemorhous The American rosefinches that form the genus ''Haemorhous'', are a group of passerine birds in the finch family Fringillidae. As the name implies ("haemo" means "blood" in Greek), various shades of red are characteristic plumage colors of this g ...
'' – contains the three North America rosefinches * '' Chloris'' – contains the six greenfinches * ''
Rhodospiza The desert finch (''Rhodospiza obsoleta''), sometimes called Lichtenstein's desert finch, is a large brown true finch found in southern Eurasia. Its taxonomy is confused, and it has formerly been placed in ''Fringilla'', '' Bucanetes'', ''Cardue ...
'' – contains a single species, the
desert finch The desert finch (''Rhodospiza obsoleta''), sometimes called Lichtenstein's desert finch, is a large brown true finch found in southern Eurasia. Its taxonomy is confused, and it has formerly been placed in ''Fringilla'', '' Bucanetes'', ''Cardu ...
* '' Rhynchostruthus'' – contains the three golden-winged grosbeaks * '' Linurgus'' – contains a single species, the
oriole finch The oriole finch (''Linurgus olivaceus'') is a small passerine bird in the finch family. It is found in Africa and is native to Burundi, Cameroon, Democratic Republic of the Congo, Equatorial Guinea, Kenya, Malawi, Nigeria, Rwanda, South Suda ...
* ''
Crithagra ''Crithagra'' is a genus of small passerine birds in the finch family (Fringillidae). They live in Africa and Arabia. The species in this genus were formerly assigned to the genus ''Serinus'', but phylogenetic analysis of mitochondrial and nucle ...
'' – contains 37 species of canaries, serins and siskins from Africa and the Arabian Peninsula * ''
Linaria ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...
'' – contains four species including the
twite The twite (''Linaria flavirostris'') is a small brown passerine bird in the finch family Fringillidae. It is similar in size and shape to a linnet, at long. It lacks the red head patch and breast shown by the linnet and the redpolls. It is bro ...
and three linnets * '' Acanthis'' – contains three redpolls * ''
Loxia The crossbill is a genus, ''Loxia'', of birds in the finch family (Fringillidae), with six species. These birds are characterised by the mandibles with crossed tips, which gives the group its English name. Adult males tend to be red or orange in ...
'' – contains six crossbills * ''
Chrysocorythus ''Chrysocorythus'' is a genus of finches in the family Fringillidae. It contains the following species: * Indonesian serin The Indonesian serin (''Chrysocorythus estherae'') is a species of finch in the family Fringillidae. It is found in In ...
'' – contains two species, the
Indonesian serin The Indonesian serin (''Chrysocorythus estherae'') is a species of finch in the family Fringillidae. It is found in Indonesia. Its natural habitats are subtropical or tropical moist montane forest and subtropical or tropical high-altitude grass ...
and the
Mindanao serin The Mindanao serin (''Chrysocorythus mindanensis'') is a species of finch in the family Fringillidae. It is found in the Philippines in the areas of Mount Katanglad and Mount Apo. Its natural habitats are subtropical or tropical moist montane f ...
* ''
Carduelis The genus ''Carduelis'' is a group of birds in the finch family Fringillidae. The genus ''Carduelis'' was introduced by the French zoologist Mathurin Jacques Brisson in 1760 by tautonomy based on Carl Linnaeus's specific epithet for the Europe ...
'' – contains three species including the
European goldfinch The European goldfinch or simply the goldfinch (''Carduelis carduelis'') is a small passerine bird in the finch family that is native to Europe, North Africa and western and central Asia. It has been introduced to other areas, including Austral ...
* ''
Serinus ''Serinus'' is a genus of small birds in the finch family Fringillidae found in Europe and Africa. The birds usually have some yellow in their plumage. The genus was introduced in 1816 by the German naturalist Carl Ludwig Koch. Its name is New L ...
'' – contains eight species including the
European serin The European serin, or simply the serin (''Serinus serinus''), is the smallest European species of the family of finches (Fringillidae) and is closely related to the Atlantic canary. Its diet consists mainly of a combination of buds and seeds. ...
* '' Spinus'' – contains 20 species including the North American goldfinches and the
Eurasian siskin The Eurasian siskin (''Spinus spinus'') is a small passerine bird in the finch family Fringillidae. It is also called the European siskin, common siskin or just siskin. Other (archaic) names include ''black-headed goldfinch'', ''barley bird'' a ...


References


Literature cited

* * {{Taxonbar, from=Q667490 Bird subfamilies Fringillidae Higher-level bird taxa restricted to the Australasia-Pacific region