Cardellina Pusilla - Wilson's Warbler - XC108402
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''Cardellina'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
birds in the
New World warbler The New World warblers or wood-warblers are a group of small, often colorful, passerine birds that make up the family Parulidae and are restricted to the New World. The family contains 120 species. They are not closely related to Old World warb ...
family Parulidae. The genus name ''Cardellina'' is a diminutive of the Italian dialect word ''Cardella'' for the
European goldfinch The European goldfinch or simply the goldfinch (''Carduelis carduelis'') is a small passerine bird in the finch Family (biology), family that is native to the Palearctic zone in Europe, northern Africa, and western Asia. It has been introduced t ...
.


Taxonomy

The genus was introduced by the French naturalist
Charles Lucien Bonaparte Charles Lucien Jules Laurent Bonaparte, 2nd Prince of Canino and Musignano (24 May 1803 – 29 July 1857) was a French naturalist and ornithology, ornithologist, and a nephew of Napoleon. Lucien and his wife had twelve children, including Cardinal ...
in 1850. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
was subsequently designated as the
red-faced warbler The red-faced warbler (''Cardellina rubrifrons'') is a species of New World warbler. Mature red-faced warblers are small birds, long. They are light gray on top with a white rump and a white underside. The face, neck, and upper breast are all b ...
. The genus originally contained one species, the red-faced warbler. A comprehensive study of the wood-warblers published in 2010 that analysed mitochondrial and nuclear DNA sequences found that the five species formed a discrete
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
, with the Wilson's and Canada warblers as early offshoots, followed by a lineage that gave rise to two branches – one leading to the red-faced and another that diverged to the red and pink-headed warblers.
Spencer Fullerton Baird Spencer Fullerton Baird (; February 3, 1823 – August 19, 1887) was an American naturalist, ornithologist, ichthyologist, Herpetology, herpetologist, and museum curator. Baird was the first curator to be named at the Smithsonian Institution. He ...
described the genus ''Ergaticus'' in 1865, using it to separate several species from what he felt was the closely related genus ''Cardellina'', and eventually subsumed into ''Cardellina'' in 2011. Prior to the creation and widespread acceptance of this genus, the
red warbler The red warbler (''Cardellina rubra'') is a small passerine bird of the New World warbler family Parulidae endemic to the highlands of Mexico, north of the Isthmus of Tehuantepec. It is closely related to, and forms a superspecies with, the p ...
and pink-headed warbler were placed in various other warbler genera, including ''Setophaga'' (with the
American redstart The American redstart (''Setophaga ruticilla'') is a New World warbler. It is unrelated to the Old World Common redstart, (common) redstart. Taxonomy The American redstart was described by Carl Linnaeus in 1758 in the 10th edition of Systema Nat ...
), ''Cardellina'' (with the
red-faced warbler The red-faced warbler (''Cardellina rubrifrons'') is a species of New World warbler. Mature red-faced warblers are small birds, long. They are light gray on top with a white rump and a white underside. The face, neck, and upper breast are all b ...
), and ''
Basileuterus ''Basileuterus'' is a genus of New World warblers, best represented in Central and South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller porti ...
'' (a widespread genus of tropical warblers) — as well as the
Old World warbler The Old World warblers are a large group of birds formerly grouped together in the bird family Sylviidae. They are not closely related to the New World warblers. The family held over 400 species in over 70 genera, and were the source of much taxo ...
genus '' Sylvia'' and the
Old World The "Old World" () is a term for Afro-Eurasia coined by Europeans after 1493, when they became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia in the Eastern Hemisphere, previously ...
tit genus ''
Parus ''Parus'' is a genus of Old World birds in the tit family Paridae. It was formerly a large genus containing most of the 50 odd species in the family Paridae. The genus was split into several resurrected genera following the publication of a deta ...
''. Recent
DNA Deoxyribonucleic acid (; DNA) is a polymer composed of two polynucleotide chains that coil around each other to form a double helix. The polymer carries genetic instructions for the development, functioning, growth and reproduction of al ...
analysis shows that ''Ergaticus'' falls comfortably within the New World warbler
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
, along with 18 other
genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
. It is closest to the genus ''Cardellina'', with which it shares a common ancestor, and slightly more distantly related to the genus '' Wilsonia''. There are two sister species, separated by the low-lying Isthmus of Tehuantepec, in the genus. The red warbler, ''C. ruber'', is found in the Mexican highlands north of the isthmus. Its three
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
, which differ slightly in appearance, are found in three disjunct populations. The pink-headed warbler, ''C. versicolor'', is found south of the isthmus, in the highlands of Chiapas, Mexico and western Guatemala. It is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
across its limited range. Though they are separated by geography and differ considerably in
plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...
, the two have sometimes been considered to be
conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organism ...
. ''Ergaticus'' is the
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
ized version of the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
''ergatikos'', meaning "willing or able to work".


List of species

The following five species are currently recognized.


Description

These are medium-sized warblers, measuring in length, and weighing ; the pink-headed warbler is, on average, slightly the heavier of the two. As adults, their overall color is red, with duller wings and tails; juveniles are tawny-brown, with slightly paler underparts. The red warbler has white or silvery-gray ear patches (the color depends on the subspecies), while the pink-headed warbler's head and chest are silvery-pink. The sexes are similar in both species. They have long, rounded wings and fairly long, rounded tails. They have small, narrow bills, with
rictal bristle The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for pecking, grasping, and holding (in probing for food, eating, manipulating and ca ...
s that extend more than halfway down their length.


Habitat and range

Both the pink-headed and red warblers are birds of highland forest. The red warbler is found from
above sea level Height above mean sea level is a measure of a location's vertical distance (height, elevation or altitude) in reference to a vertical datum based on a historic mean sea level. In geodesy, it is formalized as orthometric height. The zero level ...
, and the pink-headed warbler from .


Notes


References

Bird genera Taxa named by Charles Lucien Bonaparte {{Parulidae-stub