Canthium Umbelligerum
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''Canthium'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Rubiaceae. They are
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s and small
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s. The leaves are
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
and the stems are usually thorny.


Distribution

''Canthium'' species are predominantly found in
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
, especially in
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...
and the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
. A small number of species is found in
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
,
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
, and
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mos ...
. Only a limited number of species is found on the African continent, especially in
Southern Southern may refer to: Businesses * China Southern Airlines, airline based in Guangzhou, China * Southern Airways, defunct US airline * Southern Air, air cargo transportation company based in Norwalk, Connecticut, US * Southern Airways Express, M ...
and
East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historical ...
.


Taxonomy

''Canthium'' was named by
Jean-Baptiste Lamarck Jean-Baptiste Pierre Antoine de Monet, chevalier de Lamarck (1 August 1744 – 18 December 1829), often known simply as Lamarck (; ), was a French naturalist, biologist, academic, and soldier. He was an early proponent of the idea that biologi ...
in 1785 in Encyclopédie Méthodique. The
name A name is a term used for identification by an external observer. They can identify a class or category of things, or a single thing, either uniquely, or within a given context. The entity identified by a name is called its referent. A personal ...
is a latinisation of "kantankara", a
Malayalam Malayalam (; , ) is a Dravidian language spoken in the Indian state of Kerala and the union territories of Lakshadweep and Puducherry (Mahé district) by the Malayali people. It is one of 22 scheduled languages of India. Malayalam was des ...
name A name is a term used for identification by an external observer. They can identify a class or category of things, or a single thing, either uniquely, or within a given context. The entity identified by a name is called its referent. A personal ...
from
Kerala Kerala ( ; ) is a state on the Malabar Coast of India. It was formed on 1 November 1956, following the passage of the States Reorganisation Act, by combining Malayalam-speaking regions of the erstwhile regions of Cochin, Malabar, South ...
for ''Canthium coromandelicum''. ''Kantan'' means "shining" and ''kara'' means "a spiny shrub". The
biological type In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes the ...
for the genus consists of
specimen Specimen may refer to: Science and technology * Sample (material), a limited quantity of something which is intended to be similar to and represent a larger amount * Biological specimen or biospecimen, an organic specimen held by a biorepository ...
s originally described by
Jean-Baptiste Lamarck Jean-Baptiste Pierre Antoine de Monet, chevalier de Lamarck (1 August 1744 – 18 December 1829), often known simply as Lamarck (; ), was a French naturalist, biologist, academic, and soldier. He was an early proponent of the idea that biologi ...
as ''Canthium parviflorum''''Canthium'' In: Index Nominum Genericorum. In: Regnum Vegetabile but this species is now included in '' Canthium coromandelicum''. ''Canthium'' is a member of Vanguerieae, a
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English language, English is in the discipline of anthropology. This definition is contested, in p ...
that is
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
and easily recognized morphologically, but in which
generic Generic or generics may refer to: In business * Generic term, a common name used for a range or class of similar things not protected by trademark * Generic brand, a brand for a product that does not have an associated brand or trademark, other ...
boundaries were, for a long time, very unclear. ''Canthium'' was especially problematic, and until the 1980s, it was defined broadly and known to be polyphyletic. '' Psydrax'' was separated from it in 1985, as was '' Keetia'' in 1986. These were followed by ''
Pyrostria ''Pyrostria'' is a genus of dioecious flowering plants in the family Rubiaceae. Most of the species are endemic to Madagascar, others occur on islands in the western Indian Ocean (Mauritius, Comoros, Réunion, Rodrigues, Socotra), a few are ...
'' and ''
Multidentia ''Multidentia'' is a genus of flowering plants in the Family (biology), family Rubiaceae. Distribution ''Multidentia'' is restricted to tropical Africa with the greatest concentration of species occurring in eastern Africa. Taxonomy It was orig ...
'' in 1987. The
subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
''
Afrocanthium ''Afrocanthium'' is a genus of flowering plants in the family Rubiaceae. It consists of deciduous, unarmed trees, and shrubs. They are native to East Africa, from Sudan and Ethiopia to South Africa. Taxonomy ''Afrocanthium'' was first recogniz ...
'' was raised to generic rank in 2004, followed by '' Bullockia'' in 2009. A few species were transferred to ''Canthium'' from '' Rytigynia'' and other genera in 2004. The genus was further reduced by the transfer of species to ''
Peponidium ''Peponidium'' is a genus of flowering plants in the family Rubiaceae. The genus is endemic to Madagascar, except for '' Peponidium ovato-oblongum'' and '' Peponidium venulosum'' that are found on the Comoros, and '' Peponidium carinatum'' and ' ...
'' and ''
Pyrostria ''Pyrostria'' is a genus of dioecious flowering plants in the family Rubiaceae. Most of the species are endemic to Madagascar, others occur on islands in the western Indian Ocean (Mauritius, Comoros, Réunion, Rodrigues, Socotra), a few are ...
''. In 2016, two ''Canthium'' species endemic to the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
were transferred to a genus of their own, '' Kanapia''. The final circumscription of ''Canthium'' will remain in doubt until
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
studies achieve greater
resolution Resolution(s) may refer to: Common meanings * Resolution (debate), the statement which is debated in policy debate * Resolution (law), a written motion adopted by a deliberative body * New Year's resolution, a commitment that an individual mak ...
for the
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
containing '' Canthium coromandelicum'' and its closest relatives.


Species

,
Plants of the World Online Plants of the World Online (POWO) is an online database published by the Royal Botanic Gardens, Kew. It was launched in March 2017 with the ultimate aim being "to enable users to access information on all the world's known seed-bearing plants by ...
recognises the following species: * ''
Canthium aciculatum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
''
Ridl. Henry Nicholas Ridley CMG (1911), MA (Oxon), FRS, FLS, F.R.H.S. (10 December 1855 – 24 October 1956) was an English botanist, geologist and naturalist who lived much of his life in Singapore. He was instrumental in promoting rubber trees ...
* ''
Canthium angustifolium ''Canthium angustifolium'', with the common name narrow-leaved canthium, is native to southern tropical Asia. The large shrub is found in eastern India (including Assam), Bangladesh, and Myanmar Myanmar, ; UK pronunciations: US pronunc ...
'' Roxb. * '' Canthium arboreum'' Vidal * '' Canthium aurantiacum''
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
& L.M.Perry
* ''
Canthium berberidifolium ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Geddes * '' Canthium bipinnatum'' ( Blanco)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* ''
Canthium brunneum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espe ...
'' (
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* ''
Canthium calvum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Craib * ''
Canthium cambodianum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Pit. * ''
Canthium campanulatum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espe ...
'' Thwaites * ''
Canthium cavaleriei ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espe ...
'' H.Lév. * ''
Canthium ciliatum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espe ...
'' (D.Dietr.)
Kuntze Kuntze is a surname of German origin. People with that name include: * Carl Kuntze (1922-2006), Dutch rower who competed at the 1952 Summer Olympics * Edward J. Kuntze (1826-1870), Prussian-born American sculptor * Otto Kuntze (1843-1907), German ...
* ''
Canthium coffeoides ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Pierre ex Pit. * ''
Canthium congestiflorum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
''
Ridl. Henry Nicholas Ridley CMG (1911), MA (Oxon), FRS, FLS, F.R.H.S. (10 December 1855 – 24 October 1956) was an English botanist, geologist and naturalist who lived much of his life in Singapore. He was instrumental in promoting rubber trees ...
* ''
Canthium cordatum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espe ...
'' Dillwyn * '' Canthium coromandelicum'' ( Brum.f.) Alston * ''
Canthium depressinerve ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
''
Ridl. Henry Nicholas Ridley CMG (1911), MA (Oxon), FRS, FLS, F.R.H.S. (10 December 1855 – 24 October 1956) was an English botanist, geologist and naturalist who lived much of his life in Singapore. He was instrumental in promoting rubber trees ...
* ''
Canthium ellipticum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' (
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* ''
Canthium fenicis ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' (
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* ''
Canthium ferrugineum ''Canthium'' is a genus of flowering plants in the Family (biology), family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually Thorns, spines, and prickles, thorny. Distribution ''Canthium'' species a ...
'' Craib * ''
Canthium filipendulum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espe ...
'' Pierre ex Pit. * ''
Canthium fraternum ''Canthium'' is a genus of flowering plants in the Family (biology), family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually Thorns, spines, and prickles, thorny. Distribution ''Canthium'' species a ...
'' Miq. * ''
Canthium glaucum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Hiern * ''
Canthium gracilipes ''Canthium'' is a genus of flowering plants in the Family (biology), family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually Thorns, spines, and prickles, thorny. Distribution ''Canthium'' species a ...
'' Kurz * ''
Canthium hainanense ''Canthium'' is a genus of flowering plants in the Family (biology), family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually Thorns, spines, and prickles, thorny. Distribution ''Canthium'' species a ...
'' (
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) Lantz
* '' Canthium hirtellum''
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* ''
Canthium hispidonervosum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' ( De Wild.) C.M.Evrard * ''
Canthium horridulum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Craib * ''
Canthium horridum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Blume * ''
Canthium inerme ''Canthium inerme'' (Turkey-berry) is a tough, adaptable medium-sized tree from South Africa. It bears small edible fruits and has a variety of uses in traditional medicine. Name The Latin species name of this plant, ''"inerme"'', actually means ...
'' ( L.f.)
Kuntze Kuntze is a surname of German origin. People with that name include: * Carl Kuntze (1922-2006), Dutch rower who competed at the 1952 Summer Olympics * Edward J. Kuntze (1826-1870), Prussian-born American sculptor * Otto Kuntze (1843-1907), German ...
* ''
Canthium korthalsianum ''Canthium'' is a genus of flowering plants in the Family (biology), family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually Thorns, spines, and prickles, thorny. Distribution ''Canthium'' species a ...
'' Miq. * ''
Canthium kuntzeanum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Bridson * ''
Canthium laeve ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espe ...
'' Teijsm. & Binn. * '' Canthium lasianthoides'' Miq. * ''
Canthium libericum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Dinkl. * ''
Canthium longipes ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Geddes * ''
Canthium lucidum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' R.Br. * ''
Canthium macrocarpum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Thwaites * ''
Canthium malayense ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' K.M.Wong * ''
Canthium megacarpum ''Canthium'' is a genus of flowering plants in the Family (biology), family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually Thorns, spines, and prickles, thorny. Distribution ''Canthium'' species a ...
'' (
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
)
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* ''
Canthium megistocarpum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
''
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
& L.M.Perry
* ''
Canthium merrillianum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
''
Mabb. Professor David John Mabberley , (born May 1948) is a British-born botanist, educator and writer. Among his varied scientific interests is the taxonomy of tropical plants, especially trees of the families Labiatae, Meliaceae and Rutaceae. He ...
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Canthium merrillii ''Canthium'' is a genus of flowering plants in the Family (biology), family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually Thorns, spines, and prickles, thorny. Distribution ''Canthium'' species a ...
'' (Setch.)
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Canthium mite ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
''
Bartl. Friedrich Gottlieb Bartling (December 9, 1798 – November 20, 1875) was a German botanist who was a native of Hanover. He studied natural sciences at the University of Göttingen, and in 1818 took a botanical journey through Hungary and Croatia ...
ex
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* ''
Canthium molle ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espe ...
''
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Gamble Gambling (also known as betting or gaming) is the wagering of something of value ("the stakes") on a random event with the intent of winning something else of value, where instances of strategy are discounted. Gambling thus requires three elem ...
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Canthium moluccanum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Roxb. * ''
Canthium oblongum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' (Valeton) Kaneh. * '' Canthium oliganthum'' ( Miq.) Boerl. * '' Canthium oligocarpum'' Hiern * ''
Canthium parvifolium ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Roxb. * ''
Canthium paucinervium ''Canthium'' is a genus of flowering plants in the Family (biology), family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually Thorns, spines, and prickles, thorny. Distribution ''Canthium'' species a ...
'' (
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* ''
Canthium pedunculare ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Cav. * ''
Canthium perakanthus ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' ined. * ''
Canthium polyanthum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Miq. * ''
Canthium puberulum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espe ...
'' Thwaites ex
Hook.f. Sir Joseph Dalton Hooker (30 June 1817 – 10 December 1911) was a British botanist and explorer in the 19th century. He was a founder of geographical botany and Charles Darwin's closest friend. For twenty years he served as director of t ...
* ''
Canthium quadratum ''Canthium'' is a genus of flowering plants in the Family (biology), family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually Thorns, spines, and prickles, thorny. Distribution ''Canthium'' species a ...
'' Craib * ''
Canthium rheedei ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
''
DC. Augustin Pyramus (or Pyrame) de Candolle (, , ; 4 February 17789 September 1841) was a Swiss botanist. René Louiche Desfontaines launched de Candolle's botanical career by recommending him at a herbarium. Within a couple of years de Candolle ...
* ''
Canthium sarcocarpum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' (
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* '' Canthium sarmentosum'' Craib * ''
Canthium scabridum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
''
Ridl. Henry Nicholas Ridley CMG (1911), MA (Oxon), FRS, FLS, F.R.H.S. (10 December 1855 – 24 October 1956) was an English botanist, geologist and naturalist who lived much of his life in Singapore. He was instrumental in promoting rubber trees ...
* ''
Canthium scandens ''Canthium'' is a genus of flowering plants in the Family (biology), family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually Thorns, spines, and prickles, thorny. Distribution ''Canthium'' species a ...
'' Blume * ''
Canthium schlechterianum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
''
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
& L.M.Perry
* ''
Canthium simile ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espe ...
''
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
& Chun
* '' Canthium sordidum'' (
K.Schum. Karl Moritz Schumann (17 June 1851 – 22 March 1904) was a German botanist. Schumann was born in Görlitz. He was curator of the Botanisches Museum in Berlin-Dahlem from 1880 until 1894. He also served as the first chairman of the ''Deut ...
) Bullock
* ''
Canthium spirostylum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Miq. * ''
Canthium stellulatum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Craib * ''
Canthium strigosum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Craib * ''
Canthium strychnoides ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Craib * '' Canthium subaureum'' Craib * ''
Canthium subcapitatum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' (
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* ''
Canthium suberosum ''Canthium'' is a genus of flowering plants in the Family (biology), family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually Thorns, spines, and prickles, thorny. Distribution ''Canthium'' species a ...
'' Codd * ''
Canthium subsessilifolium ''Canthium'' is a genus of flowering plants in the Family (biology), family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually Thorns, spines, and prickles, thorny. Distribution ''Canthium'' species a ...
'' (
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* ''
Canthium tavoyanum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' (R.Parker)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* ''
Canthium travancoricum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
''
Bedd. Colonel Richard Henry Beddome (11 May 1830 – 23 February 1911) was a British military officer and naturalist in India, who became chief conservator of the Madras Forest Department. In the mid-19th century, he extensively surveyed several ...
* ''
Canthium trichophorum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Quisumb. &
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* '' Canthium umbelligerum'' Miq. * ''
Canthium vanwykii ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Tilney & Kok * ''
Canthium villarii ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Vidal * ''
Canthium violaceum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Zoll. & Moritzi


References

{{Taxonbar, from=Q3244344 Rubiaceae genera Afrotropical realm flora Indomalayan realm flora Taxa named by Jean-Baptiste Lamarck Taxonomy articles created by Polbot