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''Fraxinus americana'', the white ash or American ash, is a species of ''
ash tree ''Fraxinus'' (), commonly called ash, is a genus of flowering plants in the olive and lilac family, Oleaceae. It contains 45–65 species of usually medium to large trees, mostly deciduous, though a number of subtropical species are evergree ...
'' native to eastern and central
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. The species is native to mesophytic
hardwood Hardwood is wood from dicot trees. These are usually found in broad-leaved temperate and tropical forests. In temperate and boreal latitudes they are mostly deciduous, but in tropics and subtropics mostly evergreen. Hardwood (which comes from ...
forests from Nova Scotia west to Minnesota, south to northern Florida, and southwest to eastern Texas. Isolated populations have also been found in western Texas, Wyoming, and Colorado, and the species is reportedly
naturalized Naturalization (or naturalisation) is the legal act or process by which a non-citizen of a country may acquire citizenship or nationality of that country. It may be done automatically by a statute, i.e., without any effort on the part of the in ...
in Hawaii. There are an estimated 8 billion ash trees in the United States – the majority being the white ash trees and the green ash trees.


Characteristics

The name white ash derives from the
glaucous ''Glaucous'' (, ) is used to describe the pale grey or bluish-green appearance of the surfaces of some plants, as well as in the names of birds, such as the glaucous gull (''Larus hyperboreus''), glaucous-winged gull (''Larus glaucescens''), g ...
undersides of the leaves. It is similar in appearance to the green ash, making identification difficult. The lower sides of the leaves of white ash are lighter in color than their upper sides, and the outer surface of the twigs of white ash may be flaky or peeling. Green ash leaves are similar in color on upper and lower sides, and twigs are smoother. White Ash leaves turn yellow or red in Autumn. Despite some overlap, the two species tend to grow in different locations as well; white ash is a forest tree that commonly occurs alongside sugar maple while green ash is a pioneer species that inhabits riparian zones and disturbed areas.Common Trees of the North Carolina Piedmont
''Fraxinus americana''
/ref>New Brunswick tree and shrub
''Fraxinus americana''
The White Ash's compound leaves usually have 7 leaflets per leaf whereas other ash trees are usually more diverse. Like other species in the section ''Melioides'', ''Fraxinus americana'' is
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...
, with male and female flowers produced on separate individuals.


Cultivation and uses

White ash is one of the most used trees for everyday purposes and, to keep up with high demand, is cultivated almost everywhere possible. The wood is white and quite dense (within 20% of 670 kg/m3),White Ash
''Niche Timbers''. Retrieved on 2009-07-24.
strong, and straight-grained. Its species produces an ideal, atypical dominant excurrent structured crown. It is a traditional timber of choice for production of baseball bats and tool handles. The wood is also favorable for furniture and flooring. A study compared it to eight other different species, and it showed the highest antibacterial activity in the context of manufacturing chopping boards. Woodworkers use the timber mainly for interior uses due to high perishability in contact with ground soil. It is also used to make lobster traps. Since the 1950s, it has also become a popular choice for solid electric guitar bodies. It makes a serviceable longbow if properly worked. The wood was used in ceiling fan blades from the 1970s through the mid-1980s, though cane was sometimes simulated with plastic then. It is no longer used for ceiling fan blades in most countries. White ash is not seen in cultivation as often as green ash, due to its preference for undisturbed forest sites away from urban pollution and soil compaction. It sometimes has been planted for its consistently reliable autumn colors, which typically are bright orange and red hues as opposed to other species of ash that produce a uniform yellow color. Cultivation of white ash differs across North American continent. For example, within the City of Chicago region, 2010 statistics show most common street tree species is white ash at 6.2%. Along with third ranked green type at 4.9%, ashes combine to make up 11% percent of the city's street trees, with an overall population of 13,648,044 million standing ashes within Cook County alone. Autumn Purple, or Junginger, a wild variety of American white ash selected for its purple leaf color, was discovered by University of Wisconsin horticulturist Karl Junginger of McKay Nursery in Waterloo, Iowa. After its introduction in 1956, it quickly became the most popular and most expensive landscaping selection, surpassing the high priced ginkgo, London plane and white/burr oak. A related species, Biltmore ash, is sometimes treated as a variety of white ash. Other taxonomists argue that Biltmore ash is its own species (''Fraxinus biltmoreana''). North American native ash tree species are used by
North American frogs This is a checklist of amphibians found in Northern America, based mainly on publications by the Society for the Study of Amphibians and Reptiles. The information about range and status of almost all of these species can be found also for example i ...
as a critical food source, as leaves that fall from the trees are particularly suitable for tadpoles to feed upon in ponds (both temporary and permanent), large puddles, and other water sources. Species such as red maple, which are taking the place of ash due to the ash borer, are much less suitable for the frogs as a food source — resulting in poor frog survival rates and small frog sizes. The lack of tannins in the American ash variety makes them a good food source for frogs, but are not resistant to the ash borer. Varieties of ash from outside North America typically have much higher tannin levels and resist the borer. Maples and various non-native invasive trees, trees that are taking the place of American ash species in the North American ecosystem, typically have much higher leaf tannin levels. Ash species native to North America also provide important habitat and food for various other creatures that are native to North America.


Emerald ash borer

The emerald ash borer (''Agrilus planipennis''), also commonly known by the acronym EAB, is a green beetle native to Asia. In North America, the EAB is an invasive species, highly destructive to ash trees in its introduced range. The damage of this insect rivals that of
chestnut blight The pathogenic fungus ''Cryphonectria parasitica'' (formerly ''Endothia parasitica'') is a member of the Ascomycota (sac fungi). This necrotrophic fungus is native to East Asia and South East Asia and was introduced into Europe and North America ...
and Dutch elm disease. To put its damage in perspective, the number of chestnuts killed by the
chestnut blight The pathogenic fungus ''Cryphonectria parasitica'' (formerly ''Endothia parasitica'') is a member of the Ascomycota (sac fungi). This necrotrophic fungus is native to East Asia and South East Asia and was introduced into Europe and North America ...
was around 3.5 billion chestnut trees while there are 3.5 billion ash trees in Ohio alone. Dutch elm disease killed only 200 million elm trees while EAB threatens 7.5 billion ash trees in the United States. The insect threatens the entire North American genus ''Fraxinus''. Since its accidental introduction into the United States and Canada in the 1990s, and its subsequent detection in 2002, it has spread to eleven states and adjacent parts of Canada. It has killed at least 50 million ash trees so far and threatens to kill most of the ash trees throughout North America. White ash exhibits a little more resistance to the EAB than green ash, which has nearly no resistance; however this could also possibly be due to white ash not being used in landscaping as extensively or placed in high-stress environments. An infested tree can be recognized by premature fall color and leaf
senescence Senescence () or biological aging is the gradual deterioration of functional characteristics in living organisms. The word ''senescence'' can refer to either cellular senescence or to senescence of the whole organism. Organismal senescence inv ...
observed on affected branches between August and last week of September. Before the EAB was officially identified, such dieback symptoms were thought to have been caused by a vascular disease classified as
ash yellows ''Candidatus'' Phytoplasma fraxini is a species of phytoplasma, a specialized group of bacteria which lack a cell wall and attack the phloem of plants. This phytoplasma causes the diseases ash yellows and lilac witches' broom. Taxonomy ''Ca.' ...
. It was assumed damage was caused by the EAB taking advantage of weakened trees. Other recognizable signs regularly observed have been upper crown dieback,
epicormic shoot An epicormic shoot is a shoot growing from an epicormic bud, which lies underneath the bark of a trunk, stem, or branch of a plant. Epicormic buds lie dormant beneath the bark, their growth suppressed by hormones from active shoots higher up ...
s or sprouts, bark lesions,
frass Frass refers loosely to the more or less solid excreta of insects, and to certain other related matter. Definition and etymology ''Frass'' is an informal term and accordingly it is variously used and variously defined. It is derived from the G ...
filled larval galleries, and deformed exit holes. When the "flatheaded" beetle has fully developed, it will bore out of the tree, in this instance, with a hole in a deformed "D" shape. In past it was assumed, once any symptoms had started to show, or once a tree had become infested, it was likely too late to begin management. Today only on-site professionals diagnosing an individual tree can responsibly make such determinations.


Ash fungal disease

''
Hymenoscyphus fraxineus ''Hymenoscyphus fraxineus'' () is an ascomycete fungus that causes ash dieback, a chronic fungal disease of ash trees in Europe characterised by leaf loss and crown dieback in infected trees. The fungus was first scientifically described in 200 ...
'' is a fungal disease that attacks ash trees. The disease causes leaf loss, crown dieback, and lesions in the bark. This fungus is, for the most part, fatal, both directly and indirectly, by weakening the tree's immune system so that it is more susceptible to attacks from pests or pathogens. Because of this, younger trees are more affected, and fully mature ash trees are incapable of living long enough to reproduce. The disease has spread over most of Europe and has been confirmed in Britain, however, it has not yet been found in North America. The scale of the threat is not yet known, but is thought to be able to cause significant damage to the ash landscape of Europe and the UK.


References


External Links

* {{Authority control americana Hardwood forest plants Medicinal plants of North America Trees of the North-Central United States Plants described in 1753 Taxa named by Carl Linnaeus Trees of the Northeastern United States Trees of the South-Central United States Trees of the Southeastern United States Dioecious plants