''Canarium ovatum'', the pili ( Central Bikol and Filipino: ''pili'', ;), is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

tropical tree Tropical vegetation is any vegetation in tropical latitudes. Plant life that occurs in climates that are warm year-round is in general more biologically diverse that in other latitudes. Some tropical areas may receive abundant rain the whole ye ...

belonging to the

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

Canarium ''Canarium'' is a genus of about 100 species of tropical and subtropical trees, in the family Burseraceae. They grow naturally across tropical Africa, south and southeast Asia, Indochina, Malesia, Australia and western Pacific Islands; inclu ...

''. It is one of approximately 600 species in the family

Burseraceae The Burseraceae are a moderate-sized family of 17-19 genera and about 540 species of flowering plants. The actual numbers differ according to the time period in which a given source is written describing this family. The Burseraceae are also know ...

. ''C. ovatum'' are native to the Philippines. They are commercially cultivated in the Philippines for their edible

nuts Nut often refers to: * Nut (fruit), fruit composed of a hard shell and a seed, or a collective noun for dry and edible fruits or seeds * Nut (hardware), fastener used with a bolt Nut or Nuts may also refer to: Arts, entertainment, and media Com ...

and is believed to be indigenous to that country. The

fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...

and

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

are often vulgarized with the umbrella term of " Java almond" which mixes multiple species of the same genus, ''

''.

Description

The ''C. ovatum'' tree is a symmetrically-shaped evergreen, averaging tall with resinous wood and resistance to strong winds. It is

dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...

, with flowers borne on cymose inflorescence at the leaf

axil A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...

s of young shoots. As in

papaya The papaya (, ), papaw, () or pawpaw () is the plant species ''Carica papaya'', one of the 21 accepted species in the genus ''Carica'' of the family Caricaceae. It was first domesticated in Mesoamerica, within modern-day southern Mexico and ...

and

rambutan Rambutan (; taxonomic name: ''Nephelium lappaceum'') is a medium-sized tropical tree in the family Sapindaceae. The name also refers to the edible fruit produced by this tree. The rambutan is native to Southeast Asia. It is closely related to s ...

, functional

hermaphrodite In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many Taxonomy (biology), taxonomic groups of animals (mostly invertebrate ...

s exist in ''C. ovatum''.

Pollination Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds ...

is by insects. Flowering is frequent and

s ripen through a prolonged period of time. The ovary contains three

locule A locule (plural locules) or loculus (plural loculi) (meaning "little place" in Latin) is a small cavity or compartment within an organ or part of an organism (animal, plant, or fungus). In angiosperms (flowering plants), the term ''locule'' usu ...

s, each with two ovules; most of the time only one ovule develops. The fruit of ''C. ovatum'' is a

drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...

, long, in diameter, and weighs . The skin ( exocarp) is smooth, thin, shiny, and turns purplish black when the fruit ripens; the pulp ( mesocarp) is fibrous, fleshy, and greenish yellow in color, and the hard shell ( endocarp) within protects a normally dicotyledonous embryo. The basal end of the shell (endocarp) is pointed and the apical end is more or less blunt; between the seed and the hard shell (endocarp) is a thin, brownish, fibrous seed coat developed from the inner layer of the endocarp. This thin coat usually adheres tightly to the shell and/or the seed. Much of the kernel weight is made up of the

cotyledon A cotyledon (; ; ; , gen. (), ) is a significant part of the embryo within the seed of a plant, and is defined as "the embryonic leaf in seed-bearing plants, one or more of which are the first to appear from a germinating seed." The numb ...

s, which are about 4.1–16.6% of the whole fruit; it is composed of approximately 8%

carbohydrate In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may or ma ...

, 11.5–13.9%

protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, respo ...

, and 70% fat. Kernels from some trees may be bitter, fibrous or have a turpentine odor.

Distribution and habitat

''C. ovatums distribution range is the Philippines. ''C. ovatum'' is a tropical tree preferring deep, fertile, well drained

soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former te ...

, warm temperatures, and well distributed

rainfall Rain is water droplets that have condensed from atmospheric water vapor and then fall under gravity. Rain is a major component of the water cycle and is responsible for depositing most of the fresh water on the Earth. It provides water f ...

. It cannot tolerate the slightest frost or low temperature. Refrigeration of seeds at resulted in loss of viability after 5 days. Seed germination is highly recalcitrant, reduced from 98 to 19% after 12 weeks of storage at room temperature; seeds stored for more than 137 days did not germinate. Asexual propagations using marcotting, budding, and grafting were too inconsistent to be used in commercial production. Young shoots of ''C. ovatum'' were believed to have functional internal phloems, which rendered

bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, ...

ringing ineffective as a way of building up carbohydrate levels in the wood. Success in marcottage may be cultivar-dependent. The production standard for a mature ''C. ovatum'' tree is between of in-shell

nut Nut often refers to: * Nut (fruit), fruit composed of a hard shell and a seed, or a collective noun for dry and edible fruits or seeds * Nut (hardware), fastener used with a bolt Nut or Nuts may also refer to: Arts, entertainment, and media Com ...

, with the harvest season from May to October and peaking between June and August. There are high variations in kernel qualities and production between seedling trees. Most ''C. ovatum'' kernels tend to stick to the shell when fresh, but come off easily after being dried to 3–5% moisture ( for 27 to 28 h). Shelled nuts, with a moisture content of 2.5–4.6%, can be stored in the shade for one year without deterioration of quality.

Cultivation and uses

In the Philippines, production centers are located in the

Bicol region Bicol, known formally as the Bicol Region or colloquially as Bicolandia ( bcl, Rehiyon kan Bikol; Rinconada Bikol: ''Rehiyon ka Bikol''; Waray Sorsogon, Masbateño: ''Rehiyon san Bikol''; tl, Rehiyon ng Bikol), is an administrative region of ...

, provinces of

Sorsogon Sorsogon, officially the Province of Sorsogon (Bikol language, Bikol: ''Probinsya kan Sorsogon''; Waray language, Waray: ''Probinsya han Sorsogon''; tl, Lalawigan ng Sorsogon), is a Provinces of the Philippines, province in the Philippines locat ...

Albay Albay,