''Campanula hercegovina'', the Herzegovinian bellflower, is an

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

plant from

Bosnia and Herzegovina Bosnia and Herzegovina ( sh, / , ), abbreviated BiH () or B&H, sometimes called Bosnia–Herzegovina and often known informally as Bosnia, is a country at the crossroads of south and southeast Europe, located in the Balkans. Bosnia and H ...

. It belongs to the

Campanulaceae The family Campanulaceae (also bellflower family), of the order Asterales, contains nearly 2400 species in 84 genera of herbaceous plants, shrubs, and rarely small trees, often with milky sap. Among them are several familiar garden plants belon ...

, or bellflower,

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Description

Herzegovinian bellflower is a fragile

perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...

cushion plant A cushion plant is a compact, low-growing, mat-forming plant that is found in alpine, subalpine, arctic, or subarctic environments around the world. The term "cushion" is usually applied to woody plants that grow as spreading mats, are limited in ...

with a length of 12–20 cm, sometimes 40 cm. It has a thick, lumpy semiwoody

rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...

which can penetrate deep into the cracks of

limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms whe ...

cliffs. Its

stem Stem or STEM may refer to: Plant structures * Plant stem, a plant's aboveground axis, made of vascular tissue, off which leaves and flowers hang * Stipe (botany), a stalk to support some other structure * Stipe (mycology), the stem of a mushro ...

s are numerous, square, and highly bendable. The

leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...

of sterile individuals are gathered in tufts, ovate, pointed, with a heart-shaped base. Both sides have 2–3 teeth and stems are about three times longer than the

lamina Lamina may refer to: Science and technology * Planar lamina, a two-dimensional planar closed surface with mass and density, in mathematics * Laminar flow, (or streamline flow) occurs when a fluid flows in parallel layers, with no disruption betwee ...

. It

blossoms In botany, blossoms are the flowers of stone fruit trees (genus ''Prunus'') and of some other plants with a similar appearance that flower profusely for a period of time in spring. Colloquially, flowers of orange are referred to as such as wel ...

from June to August. The flowers are in clusters of 2–5, with 1–3 bracts. The crown is bare, light blue, split into five triangular lobes. The

fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...

is a

poricidal Dehiscence is the splitting of a mature plant structure along a built-in line of weakness to release its contents. This is common among fruits, anthers and sporangia. Sometimes this involves the complete detachment of a part; structures that op ...

capsule containing numerous

seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...

s. It opens on the side, thanks to the 3-5 holes, and the top of the fruit remains closed. The seeds are about 1 mm in length and about 0.3 mm wide. The

diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respectively ...

chromosome A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins are ...

of these bellflowers is 2''n'' = 24.

Distribution and ecology

It is endemic from the Southeast Dinaric mountains to

Albania Albania ( ; sq, Shqipëri or ), or , also or . officially the Republic of Albania ( sq, Republika e Shqipërisë), is a country in Southeastern Europe. It is located on the Adriatic and Ionian Seas within the Mediterranean Sea and shares ...

. In

, the largest local

population Population typically refers to the number of people in a single area, whether it be a city or town, region, country, continent, or the world. Governments typically quantify the size of the resident population within their jurisdiction using a ...

s are in the canyons of

Neretva The Neretva ( sr-cyrl, Неретва, ), also known as Narenta, is one of the largest rivers of the eastern part of the Adriatic basin. Four HE power-plants with large dams (higher than 150,5 metres) provide flood protection, power and water s ...

Rakitnica Rakitnica ( sr-cyrl, Ракитница) is the main tributary of the first section of the Neretva river, also called Upper Neretva ''( bs, Gornja Neretva)''. It meets Neretva from the right, flowing from north to south, between Bjelašnica and ...

Diva Grabovica Diva (; ) is the Latin word for a goddess. It has often been used to refer to a celebrated woman of outstanding talent in the world of opera, theatre, cinema, fashion and popular music. If referring to an actress, the meaning of ''diva'' is clo ...

, and Drežnica rivers, in the

Prenj Prenj ( sr-cyrl, Прењ) is a mountain range in the Dinaric Alps of southern Bosnia and Herzegovina, located in eastern Herzegovina near Mostar, Jablanica and Konjic. The highest peak is ''Zelena glava'' at . Prenj massif has at least 11 peaks ...

Čvrsnica Čvrsnica ( sr-cyrl, Чврсница, ) is a mountain in the Dinarides of Bosnia and Herzegovina, located in northern Herzegovina. Most of the mountain is located in the Herzegovina-Neretva Canton municipalities of Mostar and Jablanica, while th ...

Čabulja Čabulja ( sr-cyrl, Чабуља) is a mountain in the municipality of Mostar, Bosnia and Herzegovina. The highest point Velika Vlajna has an altitude of . See also *List of mountains in Bosnia and Herzegovina List of mountains in Bosnia and Herz ...

Piasa The Piasa ( ) or Piasa Bird is a creature from Native American mythology depicted in one of two murals painted by Native Americans on cliffsides above the Mississippi River. Its original location was at the end of a chain of limestone bluffs in ...

, and Velež mountains. This plant grows on

rocks in mountainous areas.

References

{{Taxonbar, from=Q2935461 Flora of Bosnia and Herzegovina Flora of Europe

hercegovina Herzegovina ( or ; sh-Latn-Cyrl, Hercegovina, separator=" / ", Херцеговина, ) is the southern and smaller of two main geographical region of Bosnia and Herzegovina, the other being Bosnia. It has never had strictly defined geograp ...