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''Camissoniopsis bistorta'' is a species of flowering plant in the evening primrose family known by the common names southern suncup and California suncup. It is native to southern
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
and
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
, where it grows in several types of plant community along the coast and in the coastal hills and mountain ranges. This is a hairy
annual Annual may refer to: * Annual publication, periodical publications appearing regularly once per year **Yearbook ** Literary annual * Annual plant * Annual report * Annual giving * Annual, Morocco, a settlement in northeastern Morocco * Annuals (b ...
or short-lived
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
herb In general use, herbs are a widely distributed and widespread group of plants, excluding vegetables and other plants consumed for macronutrients, with savory or aromatic properties that are used for flavoring and garnishing food, for medicinal ...
spreading from a basal rosette, with stems reaching up to 80 centimeters long. The leaves are narrow and sometimes toothed, and 1 to 12 centimeters in length. Toward the end of the spreading stems are nodding
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
s of flowers, each flower with four bright yellow petals dotted with red at their bases.


Description

This herbaceous plant grows as an
annual Annual may refer to: * Annual publication, periodical publications appearing regularly once per year **Yearbook ** Literary annual * Annual plant * Annual report * Annual giving * Annual, Morocco, a settlement in northeastern Morocco * Annuals (b ...
or short-lived
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
. It can be found in a wide range of sizes, with the young plants having a mean size of approximately . It is covered in strigose or spreading
hairs Hair is a protein filament that grows from follicles found in the dermis. Hair is one of the defining characteristics of mammals. The human body, apart from areas of glabrous skin, is covered in follicles which produce thick terminal and f ...
. The plant is occasionally simple, but most often several stems are present. The slender stems are prostrate or decumbent to relatively ascending, reaching in length, with older epidermis peeling. The leaves are alternately-arranged and measure long. Sometimes the leaves have a constriction on them, giving them a spatulate end with less hair. The basal leaves arise on petioles up to from a rosette, and are shaped narrowly elliptic. The cauline leaves tend to be (sub)
sessile Sessility, or sessile, may refer to: * Sessility (motility), organisms which are not able to move about * Sessility (botany), flowers or leaves that grow directly from the stem or peduncle of a plant * Sessility (medicine), tumors and polyps that ...
, and are generally shaped lanceolate, with the margins being minutely dentate (toothed) to roughly entire (smooth). The
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
is a
spike Spike, spikes, or spiking may refer to: Arts, entertainment, and media Books * ''The Spike'' (novel), a novel by Arnaud de Borchgrave * ''The Spike'' (book), a nonfiction book by Damien Broderick * ''The Spike'', a starship in Peter F. Hamilto ...
that is nodding in bud. The hairs on the inflorescence are short and erect. The flowers are
self-incompatible Self-incompatibility (SI) is a general name for several genetic mechanisms that prevent self-fertilization in sexually reproducing organisms, and thus encourage outcrossing and allogamy. It is contrasted with separation of sexes among individuals ...
, and open at dawn. The
hypanthium In angiosperms, a hypanthium or floral cup is a structure where basal portions of the calyx, the corolla, and the stamens form a cup-shaped tube. It is sometimes called a floral tube, a term that is also used for corolla tube and calyx tube. It ...
is long. There are 4
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
s, which are long. There are 4 yellow, fading red
petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...
s, which are long, with 1 to 2 red spots near the base. There are 8
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s, with the anthers attached at the middle. The stigma exceeds the anthers and is held well above them at anthesis. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
s measure in length, and in width. The fruits are more or less quadrangular, and are generally straight or slightly wavy and twisted. The seeds are large.


Taxonomy

The type specimen was collected by
Thomas Nuttall Thomas Nuttall (5 January 1786 – 10 September 1859) was an England, English botany, botanist and zoologist who lived and worked in America from 1808 until 1841. Nuttall was born in the village of Long Preston, near Settle, North Yorkshire, S ...
on his 1836 voyage to San Diego. The name ''bistorta'' means "twice-twisted," referring to the fact that the fruit does a double turn. Phylogenetic analysis place this species in a clade with '' Camissoniopsis hirtella'' and '' Camissoniopsis cheiranthifolia''. It hybridizes with ''C. cheiranthifolia'' subsp. ''suffruticosa''.


Distribution and habitat

This species is native to
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
in the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
and
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
in
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
. In California, it is found in the southwestern portion of the state, from
Kern KERN (1180 AM) is a commercial radio station licensed to Wasco-Greenacres, California, and serving the Bakersfield metropolitan area. The station is owned by American General Media. The radio studios and offices are in the American General Me ...
and Ventura counties south to
San Diego County San Diego County (), officially the County of San Diego, is a county in the southwestern corner of the U.S. state of California. As of the 2020 census, the population was 3,298,634, making it California's second-most populous county and the fi ...
and inland to the
Peninsular Ranges The Peninsular Ranges (also called the Lower California province) are a group of mountain ranges that stretch from Southern California to the southern tip of the Baja California peninsula; they are part of the North American Coast Ranges, which ...
and
Transverse Ranges The Transverse Ranges are a group of mountain ranges of southern California, in the Pacific Coast Ranges physiographic region in North America. The Transverse Ranges begin at the southern end of the California Coast Ranges and lie within Santa B ...
. In Baja California, this species is found from the border to inland to
Tecate Tecate () is a city in Tecate Municipality, Baja California. It is across the Mexico-US border from Tecate, California. As of 2019, the city had a population of 108,860 inhabitants, while the metropolitan area has a population of 132,406 inha ...
south to the Colonet area. Hybrids with ''Camissoniopsis cheiranthifolia'' are found on Pacific beaches between
Tijuana Tijuana ( ,"Tijuana"
(US) and
< ...
and Ensenada. This species is commonly found growing in sandy fields near the coast, but is also found in clay soils in
grassland A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natur ...
s to openings in
coastal sage scrub Coastal sage scrub, also known as coastal scrub, CSS, or soft chaparral, is a low scrubland plant community of the California coastal sage and chaparral subecoregion, found in coastal California and northwestern coastal Baja California. It is w ...
and
chaparral Chaparral ( ) is a shrubland plant community and geographical feature found primarily in the U.S. state of California, in southern Oregon, and in the northern portion of the Baja California Peninsula in Mexico. It is shaped by a Mediterranean c ...
. In some locations in San Diego County, it occurs with deceptively similar looking (when young) rare plants such as '' Chorizanthe orcuttiana'' and '' Mucronea californica''.


Gallery

File:Camissoniopsis bistorta 31765564.jpg, The flowers on their stalks File:Camissoniopsis bistorta 34776605.jpg, The characteristic rosette File:Camissoniopsis bistorta 33118028.jpg, The plant with numerous inflorescences and flowers File:Camissoniopsis bistorta 116351766.jpg, Flowers over a basal rosette of leaves File:Camissoniopsis bistorta 126379374.jpg, Flowers on ascending stems


References


External links


Jepson Manual TreatmentPhoto gallery
{{Taxonbar, from2=Q5026818, from1=Q47528209, from3=Q39309953
bistorta ''Bistorta'' is a genus of flowering plants in the family Polygonaceae. about 40 species are accepted. It has been supported as a separate clade by molecular phylogenetic analysis. ''Bistorta'' species are native throughout much of the Northern ...
Flora of Baja California Flora of California Plants described in 1840 Taxa named by Thomas Nuttall Taxa named by John Torrey Taxa named by Asa Gray