''Calophyllum'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

s in the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Calophyllaceae. They are mainly distributed in

Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...

, with some species in

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

, the Americas,

Australasia Australasia is a region that comprises Australia, New Zealand and some neighbouring islands in the Pacific Ocean. The term is used in a number of different contexts, including geopolitically, physiogeographically, philologically, and ecologica ...

, and the

Pacific Islands Collectively called the Pacific Islands, the islands in the Pacific Ocean are further categorized into three major island groups: Melanesia, Micronesia, and Polynesia. Depending on the context, the term ''Pacific Islands'' may refer to one of se ...

History

Members of the genus ''Calophyllum'' native to

Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indomalayan and Australasian realms, and also a phytogeographical floristic region in the Paleotropical Kingdom. It has been given different definitions. The ...

and

Wallacea Wallacea is a biogeographical designation for a group of mainly Indonesian islands separated by deep-water straits from the Asian and Australian continental shelves. Wallacea includes Sulawesi, the largest island in the group, as well as Lo ...

are of particular importance to traditional

shipbuilding Shipbuilding is the construction of ships and other floating vessels. It normally takes place in a specialized facility known as a shipyard. Shipbuilders, also called shipwrights, follow a specialized occupation that traces its roots to befor ...

of the larger Austronesian

outrigger ships Outrigger boats are various watercraft featuring one or more lateral support floats known as outriggers, which are fastened to one or both sides of the main hull (ship), hull. They can range from small dugout (boat), dugout canoes to large plan ...

and were carried with them in the

Austronesian expansion The Austronesian peoples, sometimes referred to as Austronesian-speaking peoples, are a large group of peoples in Taiwan, Maritime Southeast Asia, Micronesia, coastal New Guinea, Island Melanesia, Polynesia, and Madagascar that speak Austron ...

as they migrated to

Oceania Oceania (, , ) is a region, geographical region that includes Australasia, Melanesia, Micronesia, and Polynesia. Spanning the Eastern Hemisphere, Eastern and Western Hemisphere, Western hemispheres, Oceania is estimated to have a land area of ...

and

Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...

. They were comparable in importance to how

oak An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''L ...

s were in European shipbuilding and timber industries. The most notable species is the mastwood ('' Calophyllum inophyllum'') which grows readily in the sandy and rocky beaches of the island environments that the Austronesians colonized.

Description

''Calophyllum'' are

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

s or

shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...

s. They produce a colorless, white, or yellow

latex Latex is an emulsion (stable dispersion) of polymer microparticles in water. Latexes are found in nature, but synthetic latexes are common as well. In nature, latex is found as a milky fluid found in 10% of all flowering plants (angiosperms ...

. The oppositely arranged leaves have leathery blades often borne on petioles. The leaves are distinctive, with narrow parallel veins alternating with

resin In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on natu ...

canals. The

inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...

is a cyme or a thyrse of flowers that grows from the leaf axils or at the ends of branches. In the flower the

sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...

s and

petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...

s may look similar and are arranged in whorls. There are many

stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...

s. The fruit is a

drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...

with thin layers of flesh over a large seed.

Uses

Many species are used for their

wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin th ...

. Some are

hardwood Hardwood is wood from dicot trees. These are usually found in broad-leaved temperate and tropical forests. In temperate and boreal latitudes they are mostly deciduous, but in tropics and subtropics mostly evergreen. Hardwood (which comes from ...

trees that can reach 30 meters in height. They tend to grow rapidly. The outer sapwood is yellowish, yellow-brown, or orange, sometimes with a pink tinge, and the inner heartwood is light reddish to red-brown. The wood has a streaked, ribboned, or zig-zag grain. The wood has been used to build boats, flooring, and furniture, and made into

plywood Plywood is a material manufactured from thin layers or "plies" of wood veneer that are glued together with adjacent layers having their wood grain rotated up to 90 degrees to one another. It is an engineered wood from the family of manufactured ...

. ''Calophyllum'' wood may be sold under the name bitangor, and the species may be used interchangeably; one shipment may contain boards from several different species.Friday, J. B. and R. Ogoshi
Farm and Forestry Production and Marketing Profile for Tamanu (''Calophyllum inophyllum'').
In: Elevitch, C. R. (ed.) Specialty Crops for Pacific Island Agroforestry. Permanent Agriculture Resources. Holualoa, Hawaii. 2011. Plants of the genus are also known for their chemistry, with a variety of secondary metabolites isolated, such as coumarins,

xanthone Xanthone is an organic compound with the molecular formula O 6H4CO. It is a white solid. In 1939, xanthone was introduced as an insecticide and it currently finds uses as Insecticide, ovicide for codling moth eggs and as a larvicide. Xanthone ...

flavonoid Flavonoids (or bioflavonoids; from the Latin word ''flavus'', meaning yellow, their color in nature) are a class of polyphenolic secondary metabolites found in plants, and thus commonly consumed in the diets of humans. Chemically, flavonoids ...

s, and triterpenes. Compounds from the genus have been reported to have

cytotoxic Cytotoxicity is the quality of being toxic to cells. Examples of toxic agents are an immune cell or some types of venom, e.g. from the puff adder (''Bitis arietans'') or brown recluse spider (''Loxosceles reclusa''). Cell physiology Treating cells ...

, anti-

HIV The human immunodeficiency viruses (HIV) are two species of ''Lentivirus'' (a subgroup of retrovirus) that infect humans. Over time, they cause acquired immunodeficiency syndrome (AIDS), a condition in which progressive failure of the immune ...

, antisecretory, cytoprotective,

antinociceptive Nociception (also nocioception, from Latin ''nocere'' 'to harm or hurt') is the Somatosensory system, sensory nervous system's process of encoding Noxious stimulus, noxious stimuli. It deals with a series of events and processes required for an org ...

molluscicidal Molluscicides () – also known as snail baits, snail pellets, or slug pellets – are pesticides against molluscs, which are usually used in agriculture Agriculture or farming is the practice of cultivating plants and livestock. Agricul ...

, and antimicrobial properties. Some plants are used in folk medicine to treat conditions such as peptic ulcers,

tumor A neoplasm () is a type of abnormal and excessive growth of tissue. The process that occurs to form or produce a neoplasm is called neoplasia. The growth of a neoplasm is uncoordinated with that of the normal surrounding tissue, and persists ...

infection An infection is the invasion of tissues by pathogens, their multiplication, and the reaction of host tissues to the infectious agent and the toxins they produce. An infectious disease, also known as a transmissible disease or communicable dise ...

pain Pain is a distressing feeling often caused by intense or damaging stimuli. The International Association for the Study of Pain defines pain as "an unpleasant sensory and emotional experience associated with, or resembling that associated with, ...

, and

inflammation Inflammation (from la, wikt:en:inflammatio#Latin, inflammatio) is part of the complex biological response of body tissues to harmful stimuli, such as pathogens, damaged cells, or Irritation, irritants, and is a protective response involving im ...

. ''C. inophyllum'' is the source of tamanu oil, a greenish, nutty-scented oil of commercial value. It has been used as massage oil, topical medicine,

lamp oil Kerosene, paraffin, or lamp oil is a combustible hydrocarbon liquid which is derived from petroleum. It is widely used as a fuel in aviation as well as households. Its name derives from el, κηρός (''keros'') meaning "wax", and was regis ...

, and

waterproofing Waterproofing is the process of making an object or structure waterproof or water-resistant so that it remains relatively unaffected by water or resisting the ingress of water under specified conditions. Such items may be used in wet environme ...

, and is still used in

cosmetics Cosmetics are constituted mixtures of chemical compounds derived from either natural sources, or synthetically created ones. Cosmetics have various purposes. Those designed for personal care and skin care can be used to cleanse or protect ...

Tacamahac Tacamahac is the name of medicinal resins, now little used, obtained from several plant sources including '' Calophyllum tacamahaca'' and ''Calophyllum inophyllum''. The word has sometimes been regarded, apparently wrongly, as a synonym of balm of ...

is the resin of the tree. This species is also cultivated for its wood and planted in coastal landscaping as a

windbreak A windbreak (shelterbelt) is a planting usually made up of one or more rows of trees or shrubs planted in such a manner as to provide shelter from the wind and to protect soil from erosion. They are commonly planted in hedgerows around the edges ...

and for

erosion Erosion is the action of surface processes (such as water flow or wind) that removes soil, rock, or dissolved material from one location on the Earth's crust, and then transports it to another location where it is deposited. Erosion is distin ...

control.

Symbolism

A stylized ''Calophyllum'' is featured on the national

coat of arms of Nauru The design of the coat of arms of the Republic of Nauru originated in 1968 following the declaration of independence, and it began to be used officially in the early 1970s. Its shield is divided and separated in the middle. In the upper section ...

Diversity

There are approximately 187 species in the genus. Species include:

Gallery

Calophyllum caledonicum.jpg, ''

Calophyllum caledonicum ''Calophyllum'' is a genus of tropical flowering plants in the family Calophyllaceae. They are mainly distributed in Asia, with some species in Africa, the Americas, Australasia, and the Pacific Islands. History Members of the genus ''Calophyllu ...

'' File:Calophyllum brasiliense plantation.jpg, ''

Calophyllum brasiliense ''Calophyllum brasiliense'' (guanandi) is a species of plant in the family ''Calophyllaceae''. It is native to subtropical and tropical regions of Mexico, Central America, South America and the Caribbean. Description It is an evergreen tree grow ...

'' plantation Calophyllum antillanum (Palo María) picture 1.png, '' Calophyllum antillanum''

References

Bibliography

* * * * {{Taxonbar, from=Q2697391 Malpighiales genera National symbols of Nauru