Calophyllum Apetalum
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''Calophyllum'' is a genus of tropical flowering plants in the family Calophyllaceae. They are mainly distributed in Asia, with some species in Africa, the Americas, Australasia, and the
Pacific Islands Collectively called the Pacific Islands, the islands in the Pacific Ocean are further categorized into three major island groups: Melanesia, Micronesia, and Polynesia. Depending on the context, the term ''Pacific Islands'' may refer to one of se ...
.


History

Members of the genus ''Calophyllum'' native to Malesia and Wallacea are of particular importance to traditional shipbuilding of the larger
Austronesian Austronesian may refer to: *The Austronesian languages *The historical Austronesian peoples The Austronesian peoples, sometimes referred to as Austronesian-speaking peoples, are a large group of peoples in Taiwan, Maritime Southeast Asia, M ...
outrigger ships Outrigger boats are various watercraft featuring one or more lateral support floats known as outriggers, which are fastened to one or both sides of the main hull (ship), hull. They can range from small dugout (boat), dugout canoes to large plan ...
and were carried with them in the Austronesian expansion as they migrated to Oceania and Madagascar. They were comparable in importance to how oaks were in European shipbuilding and timber industries. The most notable species is the mastwood ('' Calophyllum inophyllum'') which grows readily in the sandy and rocky beaches of the island environments that the Austronesians colonized.


Description

''Calophyllum'' are trees or
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s. They produce a colorless, white, or yellow latex. The oppositely arranged leaves have leathery blades often borne on petioles. The leaves are distinctive, with narrow parallel veins alternating with resin canals. The inflorescence is a cyme or a thyrse of flowers that grows from the leaf axils or at the ends of branches. In the flower the sepals and
petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...
s may look similar and are arranged in whorls. There are many
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s. The fruit is a
drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...
with thin layers of flesh over a large seed.


Uses

Many species are used for their wood. Some are
hardwood Hardwood is wood from dicot trees. These are usually found in broad-leaved temperate and tropical forests. In temperate and boreal latitudes they are mostly deciduous, but in tropics and subtropics mostly evergreen. Hardwood (which comes from ...
trees that can reach 30 meters in height. They tend to grow rapidly. The outer sapwood is yellowish, yellow-brown, or orange, sometimes with a pink tinge, and the inner heartwood is light reddish to red-brown. The wood has a streaked, ribboned, or zig-zag grain. The wood has been used to build boats, flooring, and furniture, and made into
plywood Plywood is a material manufactured from thin layers or "plies" of wood veneer that are glued together with adjacent layers having their wood grain rotated up to 90 degrees to one another. It is an engineered wood from the family of manufactured ...
. ''Calophyllum'' wood may be sold under the name bitangor, and the species may be used interchangeably; one shipment may contain boards from several different species.Friday, J. B. and R. Ogoshi
Farm and Forestry Production and Marketing Profile for Tamanu (''Calophyllum inophyllum'').
In: Elevitch, C. R. (ed.) Specialty Crops for Pacific Island Agroforestry. Permanent Agriculture Resources. Holualoa, Hawaii. 2011.
Plants of the genus are also known for their chemistry, with a variety of secondary metabolites isolated, such as coumarins,
xanthone Xanthone is an organic compound with the molecular formula O 6H4CO. It is a white solid. In 1939, xanthone was introduced as an insecticide and it currently finds uses as Insecticide, ovicide for codling moth eggs and as a larvicide. Xanthone ...
s,
flavonoid Flavonoids (or bioflavonoids; from the Latin word ''flavus'', meaning yellow, their color in nature) are a class of polyphenolic secondary metabolites found in plants, and thus commonly consumed in the diets of humans. Chemically, flavonoids ...
s, and triterpenes. Compounds from the genus have been reported to have
cytotoxic Cytotoxicity is the quality of being toxic to cells. Examples of toxic agents are an immune cell or some types of venom, e.g. from the puff adder (''Bitis arietans'') or brown recluse spider (''Loxosceles reclusa''). Cell physiology Treating cells ...
, anti- HIV, antisecretory, cytoprotective,
antinociceptive Nociception (also nocioception, from Latin ''nocere'' 'to harm or hurt') is the Somatosensory system, sensory nervous system's process of encoding Noxious stimulus, noxious stimuli. It deals with a series of events and processes required for an org ...
,
molluscicidal Molluscicides () – also known as snail baits, snail pellets, or slug pellets – are pesticides against molluscs, which are usually used in agriculture Agriculture or farming is the practice of cultivating plants and livestock. Agricul ...
, and antimicrobial properties. Some plants are used in folk medicine to treat conditions such as peptic ulcers, tumors, infections,
pain Pain is a distressing feeling often caused by intense or damaging stimuli. The International Association for the Study of Pain defines pain as "an unpleasant sensory and emotional experience associated with, or resembling that associated with, ...
, and inflammation. ''C. inophyllum'' is the source of tamanu oil, a greenish, nutty-scented oil of commercial value. It has been used as massage oil, topical medicine,
lamp oil Kerosene, paraffin, or lamp oil is a combustible hydrocarbon liquid which is derived from petroleum. It is widely used as a fuel in aviation as well as households. Its name derives from el, κηρός (''keros'') meaning "wax", and was regis ...
, and waterproofing, and is still used in cosmetics.
Tacamahac Tacamahac is the name of medicinal resins, now little used, obtained from several plant sources including '' Calophyllum tacamahaca'' and ''Calophyllum inophyllum''. The word has sometimes been regarded, apparently wrongly, as a synonym of balm of ...
is the resin of the tree. This species is also cultivated for its wood and planted in coastal landscaping as a windbreak and for erosion control.


Symbolism

A stylized ''Calophyllum'' is featured on the national
coat of arms of Nauru The design of the coat of arms of the Republic of Nauru originated in 1968 following the declaration of independence, and it began to be used officially in the early 1970s. Its shield is divided and separated in the middle. In the upper section ...
.


Diversity

There are approximately 187 species in the genus. Species include:


Gallery

Calophyllum caledonicum.jpg, ''
Calophyllum caledonicum ''Calophyllum'' is a genus of tropical flowering plants in the family Calophyllaceae. They are mainly distributed in Asia, with some species in Africa, the Americas, Australasia, and the Pacific Islands. History Members of the genus ''Calophyllu ...
'' File:Calophyllum brasiliense plantation.jpg, ''
Calophyllum brasiliense ''Calophyllum brasiliense'' (guanandi) is a species of plant in the family ''Calophyllaceae''. It is native to subtropical and tropical regions of Mexico, Central America, South America and the Caribbean. Description It is an evergreen tree grow ...
'' plantation Calophyllum antillanum (Palo María) picture 1.png, '' Calophyllum antillanum''


See also

* Domesticated plants and animals of Austronesia


References


Bibliography

* * * * {{Taxonbar, from=Q2697391 Malpighiales genera National symbols of Nauru