Calidris Alba
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The sanderling (''Calidris alba'') is a small wading bird. The name derives from
Old English Old English (, ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages. It was brought to Great Britain by Anglo-Saxon settlement of Britain, Anglo ...
''sand-yrðling'', "sand-ploughman". The genus name is from Ancient Greek ''kalidris'' or ''skalidris'', a term used by Aristotle for some grey-coloured waterside birds. The specific, ''alba'', is Latin for "white". It is a circumpolar Arctic breeder, and is a long-distance migrant, wintering south to South America, South Europe, Africa, and
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
. It is highly gregarious in winter, sometimes forming large flocks on coastal mudflats or sandy beaches. It is somewhat unlike other sandpipers in appearance, which has led to the suggestion that it should be placed into a monotypic genus ''Crocethia''. A more recent review (Thomas ''et al.'', 2004) indicates, however, that the sanderling is a fairly typical " stint" or small sandpiper and should be separated from the large knots with its closest relatives in a distinct genus. This bird is similar in size to a
dunlin The dunlin (''Calidris alpina'') is a small wader, formerly sometimes separated with the other "stints" in the genus ''Erolia''. The English name is a dialect form of "dunling", first recorded in 1531–1532. It derives from ''dun'', "dull brown ...
, but stouter, with a thick bill. It shows a strong white wingbar in flight, and runs along the sandy beaches it prefers with a characteristic "bicycling" action of its legs, stopping frequently to pick small food items. It eats small
crab Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen) ( el, βραχύς , translit=brachys = short, / = tail), usually hidden entirely under the thorax. They live in all the ...
s and other small invertebrates. In spring, birds migrating north from South America consume large numbers of horseshoe crab eggs in the Delaware Bay area. In spring, the birds arrive on the High Arctic breeding grounds (see map), where they lay 3–4 eggs in a ground scrape. On the nesting grounds, these birds mainly eat insects and some plant material. The sanderling was described by the German naturalist Peter Simon Pallas in 1764 and given the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Trynga alba''. The sanderling is one of the species to which the ''Agreement on the Conservation of African-Eurasian Migratory Waterbirds'' ( AEWA) applies.


Description

The sanderling is a small plump sandpiper, in length. Its weight ranges from . The winter bird is very pale, almost white apart from a dark shoulder patch. This is the source of the specific name, ''alba'', which is the Latin for "white". Later in the summer, the face and throat become brick-red. The juvenile bird is spangled black and white, and shows much more contrast than the adult. If its size is misjudged, a sanderling in breeding plumage can be mistaken for some varieties of stint, or a sanderling in winter plumage can be mistaken for a
dunlin The dunlin (''Calidris alpina'') is a small wader, formerly sometimes separated with the other "stints" in the genus ''Erolia''. The English name is a dialect form of "dunling", first recorded in 1531–1532. It derives from ''dun'', "dull brown ...
or red knot. It can be told from other small wading birds, given good views, by its lack of a hind toe. Its behavior is also distinctive.


Distribution, habitat and migration

The sanderling breeds in the High Arctic areas of
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
, Europe and Asia. In North America, it breeds in the Canadian Arctic Archipelago,
Nunavut Nunavut ( , ; iu, ᓄᓇᕗᑦ , ; ) is the largest and northernmost Provinces and territories of Canada#Territories, territory of Canada. It was separated officially from the Northwest Territories on April 1, 1999, via the ''Nunavut Act'' ...
, Greenland (and to a lesser extent Alaska). In Eurasia, it breeds in
Spitsbergen Spitsbergen (; formerly known as West Spitsbergen; Norwegian: ''Vest Spitsbergen'' or ''Vestspitsbergen'' , also sometimes spelled Spitzbergen) is the largest and the only permanently populated island of the Svalbard archipelago in northern Norw ...
and areas of northern Russia from the Taymyr Peninsula to the New Siberian Islands. In the northern winter, it has a nearly
cosmopolitan distribution In biogeography, cosmopolitan distribution is the term for the range of a taxon that extends across all or most of the world in appropriate habitats. Such a taxon, usually a species, is said to exhibit cosmopolitanism or cosmopolitism. The ext ...
across the world's marine coasts. It is a complete migrant, travelling between from its breeding grounds to its wintering sites. Birds that travel further also arrive later and leave sooner. Most adults leave the breeding grounds in July and early August, whereas juvenile birds leave in late August and early September. The northward migration begins in March at the southern end of their winter distribution. The breeding habitat of the sanderling is coastal tundra north of July isotherm. The species typically chooses nesting sites on dry stony areas near wet areas, from above sea level to . During the winter and its migration, it is most commonly found on coastal sandy beaches, but also occurs on tidal sand flats, mud flats and the shores of lakes and rivers. More infrequently, it may occur on rocky shores.


Subspecies

The sanderling consists of two subspecies: *''C. a. alba'', ( Pallas, 1764), breeds on
Ellesmere Island Ellesmere Island ( iu, script=Latn, Umingmak Nuna, lit=land of muskoxen; french: île d'Ellesmere) is Canada's northernmost and List of Canadian islands by area, third largest island, and the List of islands by area, tenth largest in the world. ...
, north & east Greenland,
Svalbard Svalbard ( , ), also known as Spitsbergen, or Spitzbergen, is a Norwegian archipelago in the Arctic Ocean. North of mainland Europe, it is about midway between the northern coast of Norway and the North Pole. The islands of the group range ...
, Franz Josef Land and the Taymyr Peninsula *''C. a. rubida'', (
Gmelin Gmelin may refer to: * Gmelin's test, a chemical test * Gmelin database, a German handbook/encyclopedia of inorganic compounds initiated by Leopold Gmelin People * Carl Christian Gmelin (1762–1837), German botanist, author of ''Flora Badensis ...
, 1789)
, breeds in northeast Siberia, Alaska and northern Canada


Behaviour


Feeding behaviour

Sanderlings feed on invertebrate prey buried in the sand in the upper
intertidal zone The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of habitats with various species o ...
. In North America, this diet largely consists of the isopods '' Excirolana linguifrons'', ''
Excirolana kincaidii ''Excirolana'' is a genus of isopods in the family Cirolanidae. There are about 15 described species in ''Excirolana''. Species * '' Excirolana affinis'' (Jones, 1971) * '' Excirolana argentinae'' (Giambiagi, 1931) * '' Excirolana armata'' (Dana, ...
'', and the mole crab, ''
Emerita analoga ''Emerita analoga'', the Pacific sand crab or Pacific mole crab, is a species of small, sand-burrowing decapod crustacean found living in the sand along the temperate western coasts of North and South America. It is found on exposed sandy beache ...
''. When the tide is out, these crustaceans live in burrows some way beneath the surface. When the tide comes in, they move into the upper layers of sand and feed on the plankton and
detritus In biology, detritus () is dead particulate organic material, as distinguished from dissolved organic material. Detritus typically includes the bodies or fragments of bodies of dead organisms, and fecal material. Detritus typically hosts commun ...
that washes over them with each wave. They then burrow rapidly down again as the water retreats. They leave no marks on the surface, so the sanderlings hunt for them by plunging their beaks into the sand at random, consuming whatever they find. Their bills can penetrate only and as the water swirls around and retreats, the sand is softer; this makes it easier for the birds' beaks to penetrate further. In the spring, when much breeding activity is taking place in the
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
community, there may be as many as 4000 invertebrates per square metre, but their average size is smaller than later in the year. The birds appear to rush madly around at the edge of the surf, but in reality they are maximising their chances of catching as many prey animals as possible when they are at their most vulnerable near the surface.


Breeding behaviour

At breeding time sanderlings are
territorial A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...
, with the male aggressively defending its territory. They may either form monogamous pairs or polyandrous (one female and two male) pairings.


Gallery

File:Sanderling Dan Pancamo.jpg, Sanderling at High Island, Texas File:SANDERLING.jpg, A Sanderling in
Cagayan de Oro Cagayan ( ), officially the Province of Cagayan ( ilo, Probinsia ti Cagayan; ibg, Provinsiya na Cagayan; itv, Provinsiya ya Cagayan; fil, Lalawigan ng Cagayan), is a province in the Philippines located in the Cagayan Valley region, covering ...
, Philippines File:SanderlingsRunningFromWavesCropped.jpg, A flock displaying their distinctive behavior of running with the ebb and flow of waves (while feeding).
Willapa Bay Willapa Bay () is a bay located on the southwest Pacific coast of Washington state in the United States. The Long Beach Peninsula separates Willapa Bay from the greater expanse of the Pacific Ocean. With over of surface area Willapa Bay is the ...
, near
Tokeland, Washington Tokeland is a census-designated place (CDP) in Pacific County, Washington, United States. The population was 151 at the 2010 census, a sharp decrease from the 194 at the 2000 census. The town was named after Chief Toke, an Indian chief of the ...
. File:OneLeggedSanderling.jpg, Resting at the Monterey Bay Aquarium aviary. File:Sanderling-floreana.jpg, Floreana Island, Galapagos Islands File:Sanderling (with tag).jpg, A banded bird File:Calidris-alba-001.jpg, Running File:Sanderling from the Crossley ID Guide Britain and Ireland.jpg, ID composite File:Sanderling-11MAY2017.JPG, Sanderling in Monterey, California File:Sanderling (Calidris alba).jpg, Amrum.jpg, Sanderling on the island of Amrum, Schleswig-Holstein File:Sanderlings (72649).jpg, Sanderlings feeding in Quogue, New York


Notes


Footnotes


References

*


External links


Sanderling Species text in The Atlas of Southern African Birds
*

— Cornell Lab of Ornithology

— USGS Patuxent Bird Identification InfoCenter * * * * * * * * {{Authority control Calidris Erolia Sandpipers Birds of the Arctic Birds of the Dominican Republic Birds of the Caribbean Cosmopolitan birds Birds described in 1764 Taxa named by Peter Simon Pallas Holarctic birds