''Calamus australis'', commonly known as wait-a-while, hairy mary or lawyer cane, is a plant in the palm family Arecaceae which is

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...

to the rainforests of north east

Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , establishe ...

, Australia. Like other species in the

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

Calamus Calamus may refer to: Botany and zoology * ''Calamus'' (fish), a genus of fish in the family Sparidae * ''Calamus'' (palm), a genus of rattan palms * Calamus, the hollow shaft of a feather, also known as the quill * '' Acorus calamus'', the swe ...

'', this is a climbing plant with a very long and flexible stem. It uses sharp strong hooks on the

fronds A frond is a large, divided leaf. In both common usage and botanical nomenclature, the leaves of ferns are referred to as fronds and some botanists restrict the term to this group. Other botanists allow the term frond to also apply to the lar ...

and

tendrils In botany, a tendril is a specialized stem, leaf or petiole with a threadlike shape used by climbing plants for support and attachment, as well as cellular invasion by parasitic plants such as ''Cuscuta''. There are many plants that have tendr ...

to attach itself to other vegetation, such as taller established trees, thus gaining support that enables it to grow higher towards the

canopy Canopy may refer to: Plants * Canopy (biology), aboveground portion of plant community or crop (including forests) * Canopy (grape), aboveground portion of grapes Religion and ceremonies * Baldachin or canopy of state, typically placed over an ...

. This species is very similar to '' C. radicalis'', with which it coexists, but is smaller in almost all respects.

Description

Stems may reach lengths of up to and a diameter of . The leaves (or

) are produced on the youngest part of the stem, and the older sections are bare as the leaves age and fall off (exactly the same process as the more familiar upright palm with a stout stem). The older, leafless part of the stem of ''Calamus australis'' is smooth and glossy green. The leaves have a long

leaf sheath A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, ste ...

clasping the stem, which is densely covered with needle-like spines measuring up to long, but usually less.

Tendril In botany, a tendril is a specialized stem, leaf or petiole with a threadlike shape used by climbing plants for support and attachment, as well as cellular invasion by parasitic plants such as '' Cuscuta''. There are many plants that have tend ...

s, measuring several metres in length and armed with many stout, recurved barbs are produced from the leaf sheath. The leaf itself is

pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...

, up to in length, and has around 20 to 25 pairs of leaflets. Leaves are armed with stout recurved barbs on the lower surfaces of the

rachis In biology, a rachis (from the grc, ῥάχις [], "backbone, spine") is a main axis or "shaft". In zoology and microbiology In vertebrates, ''rachis'' can refer to the series of articulated vertebrae, which encase the spinal cord. In this c ...

. Unlike its very similar sister species ''C. radicalis'', it does not have spines on the leaflets themselves. The overall outline of the leaf is

elliptic In mathematics, an ellipse is a plane curve surrounding two focal points, such that for all points on the curve, the sum of the two distances to the focal points is a constant. It generalizes a circle, which is the special type of ellipse in ...

and the longest leaflets are at the midpoint, measuring about long by wide. The

inflorescence An inflorescence is a group or cluster of flowers arranged on a Plant stem, stem that is composed of a main branch or a complicated arrangement of branches. Morphology (biology), Morphologically, it is the modified part of the shoot of sperma ...

s are pendulous panicles measuring up to in length, carrying either

staminate The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...

(functionally male) or

pistillate Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl (botany), whorl of a flower; it consists of (one or m ...

(functionally female) flowers. Fruits are a white or cream drupe, around diameter, with a scaly exterior. They contain a single round seed which is surrounded by a thin layer of soft edible flesh.

Taxonomy

''Calamus australis'' was first described 1838 by the

German German(s) may refer to: * Germany (of or related to) ** Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ge ...

botanist

Carl Friedrich Philipp von Martius Carl Friedrich Philipp (Karl Friedrich Philipp) von Martius (17 April 1794 – 13 December 1868) was a German botanist and explorer. Life Martius was born at Erlangen, the son of Prof Ernst Wilhelm Martius, court apothecary. He graduated PhD ...

in volume 3 of his highly regarded work ''

Historia naturalis palmarum ''Historia naturalis palmarum: opus tripartitum'' ("Natural History of Palms, a work in three volumes") is a highly illustrated, three-volume botany, botanical book of Arecaceae, palms (Arecaceae) by German botanist Carl Friedrich Philipp von Ma ...

''.

Distribution and habitat

This is the most widespread of the eight Australian species of ''Calamus''. It is

tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...

areas of Queensland, from the tip of

Cape York Peninsula Cape York Peninsula is a large peninsula located in Far North Queensland, Australia. It is the largest unspoiled wilderness in northern Australia.Mittermeier, R.E. et al. (2002). Wilderness: Earth’s last wild places. Mexico City: Agrupació ...

south to near Rockhampton on the

Tropic of Capricorn The Tropic of Capricorn (or the Southern Tropic) is the circle of latitude that contains the subsolar point at the December (or southern) solstice. It is thus the southernmost latitude where the Sun can be seen directly overhead. It also reac ...

, and from sea level to about . It grows in

gallery forest A gallery forest is one formed as a corridor along rivers or wetlands, projecting into landscapes that are otherwise only sparsely treed such as savannas, grasslands, or deserts. The gallery forest maintains a more temperate microclimate above th ...

and well developed rainforest.

Ecology

Fruits of this species are eaten by

fruit dove The fruit doves, also known as fruit pigeons, are a genus (''Ptilinopus'') of birds in the pigeon and dove family (Columbidae). These colourful, frugivorous doves are found in forests and woodlands in Southeast Asia and Oceania. It is a large gen ...

s (genus ''

Ptilinopus The fruit doves, also known as fruit pigeons, are a genus (''Ptilinopus'') of birds in the pigeon and dove family (Columbidae). These colourful, frugivorous doves are found in forests and woodlands in Southeast Asia and Oceania. It is a large gen ...

'') and cassowaries (Casuarius casuarius).

Conservation

This species is listed by the Queensland Department of Environment and Science as

least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...

. , it has not been assessed by the IUCN.

Uses

Indigenous Australians Indigenous Australians or Australian First Nations are people with familial heritage from, and membership in, the ethnic groups that lived in Australia before British colonisation. They consist of two distinct groups: the Aboriginal peoples ...

had many used for this plant and its sister species. The young shoots were eaten, along with the fruits. The long flexible stem was used to construct shelters, make axe handles, fish traps, snares, and waist straps for climbing trees, and the thorny tendrils were used to catch fish.

Gallery

File:Calamus-australis-2-SF21348-03.jpg, Rachis with spines File:Calamus-australis-3-SF21348-04.jpg, Fruit File:Calamus-australis-4-SF21348-01.jpg, Foliage and stem File:Calamus-australis-SF22246-03.jpg, Tendril File:Calamus-australis-SF22180-03.jpg, Underside of leaf, showing barbs on rachis

References

External links

* *
View a map
of historical sightings of this species at the

Australasian Virtual Herbarium The ''Australasian Virtual Herbarium'' (AVH) is an online resource that allows access to plant specimen data held by various Australian and New Zealand herbaria. It is part of the Atlas of Living Australia (ALA), and was formed by the amalgama ...

View observations
of this species on

iNaturalist iNaturalist is a social network of naturalists, citizen scientists, and biologists built on the concept of mapping and sharing observations of biodiversity across the globe. iNaturalist may be accessed via its website or from its mobile applic ...

View images
of this species on Flickriver australis Endemic flora of Queensland Taxa named by Carl Friedrich Philipp von Martius Plants described in 1838 Vines {{Taxonbar , from=Q5018495