Calamosaurus
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''Calamosaurus'' (meaning "
reed Reed or Reeds may refer to: Science, technology, biology, and medicine * Reed bird (disambiguation) * Reed pen, writing implement in use since ancient times * Reed (plant), one of several tall, grass-like wetland plants of the order Poales * Re ...
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
") was a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of small
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
from the
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
-age
Lower Cretaceous Lower may refer to: *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Lower Wick is a small hamlet located in the county of Gloucestershire, England. It is situated about five miles south west of Dursley, eight ...
Wessex Formation The Wessex Formation is a fossil-rich English geological formation that dates from the Berriasian to Barremian stages (about 145–125 million years ago) of the Early Cretaceous. It forms part of the Wealden Group and underlies the younger Vecti ...
of the
Isle of Wight The Isle of Wight ( ) is a county in the English Channel, off the coast of Hampshire, from which it is separated by the Solent. It is the largest and second-most populous island of England. Referred to as 'The Island' by residents, the Isle of ...
,
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
. It is based on two
cervical In anatomy, cervical is an adjective that has two meanings: # of or pertaining to any neck. # of or pertaining to the female cervix: i.e., the ''neck'' of the uterus. *Commonly used medical phrases involving the neck are **cervical collar **cervic ...
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
e (BMNH R901), collected by Reverend William Fox.


History and taxonomy

Richard Lydekker Richard Lydekker (; 25 July 1849 – 16 April 1915) was an English naturalist, geologist and writer of numerous books on natural history. Biography Richard Lydekker was born at Tavistock Square in London. His father was Gerard Wolfe Lydekker, ...
came across these bones when cataloguing the
Fox Foxes are small to medium-sized, omnivorous mammals belonging to several genera of the family Canidae. They have a flattened skull, upright, triangular ears, a pointed, slightly upturned snout, and a long bushy tail (or ''brush''). Twelve sp ...
collection and named them ''
Calamospondylus ''Calamospondylus'' (meaning "quill vertebrae") is a genus of theropod dinosaur. It lived during the Early Cretaceous and its fossils were found on the Isle of Wight in southern England. The type species is ''C. oweni''. The holotype was collec ...
foxi'', noting their similarity to those of ''
Coelurus ''Coelurus'' ( ) is a genus of coelurosaurian dinosaur from the Late Jurassic period (mid-late Kimmeridgian faunal stage, 155–152 million years ago). The name means "hollow tail", referring to its hollow tail vertebrae (Greek ÎºÎ¿á¿–Î»Î¿Ï ...
''.Lydekker. R. (1889). On a coelurid dinosaur from the Wealden. ''Geological Magazine'' 6:119-121. Unfortunately, ''Calamospondylus'' had already been coined in 1866 (ironically by Reverend Fox himself, the very man honored in Lydekker's species name).Fox, W.D. in Anonymous. (1866) Another Wealden reptile. ''Athenaeum'' 2014:740. Lydekker renamed it in 1891 to its present title.Lydekker. R. (1891). On certain ornithosaurian and dinosaurian remains. ''Quarterly Journal of the Geological Society of London'' 47:41-44. He also at this time provisionally referred to it the right
tibia The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...
NHMUK R.186, which was identified by Naish et al. (2001) as belonging to a basal
coelurosauria Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyrann ...
n like a
compsognathid Compsognathidae is a family of coelurosaurian theropod dinosaurs. Compsognathids were small carnivores, generally conservative in form, hailing from the Jurassic and Cretaceous Periods. The bird-like features of these species, along with other ...
, and has recently been referred to
Ornithomimosauria Ornithomimosauria ("bird-mimic lizards") are theropod dinosaurs which bore a superficial resemblance to the modern-day ostrich. They were fast, omnivorous or herbivorous dinosaurs from the Cretaceous Period (geology), Period of Laurasia (now Asia ...
by Allain et al. (2014).Naish, D., Hutt, S., and Martill, D.M. (2001). Saurischian dinosaurs 2: theropods. In: Martill, D.M., and Naish, D. (eds.). ''Dinosaurs of the Isle of Wight.'' The Palaeontological Association:London, 242-309. Because of its sparse remains, it has received little attention. Often, it has been synonymized with ''Calamospondylus'' as part of a long, confusing taxonomic tangle,Swinton, W.E. (1936). The dinosaurs of the Isle of Wight. ''Proceedings of the Geologists' Association'' 47(3):204-220.Romer, A.S. (1956). ''Osteology of the Reptiles.'' University of Chicago Press:Chicago, 1-772. Steel, R. (1970). Part 14. Saurischia. ''Handbuch der Paläoherpetologie/Encyclopedia of Paleoherpetology.'' Part 14. Gustav Fischer Verlag:Stuttgart, 1-87. although there is no comparable material between the two genera. Modern reviews have regarded it as a dubious theropod,Norman, D.B. (1990). Problematic theropoda: "coelurosaurs". In: Weishampel, D.B., Dodson, P., and Osmólska, H. (eds.). ''The Dinosauria''. University of California Press:Berkeley, 280-305. .Holtz Jr., T.R., Molnar, R.E., and Currie, P.J. (2004). Basal Tetanurae. In: Weishampel, D.B., Dodson, P., and Osmólska, H. (eds.). ''The Dinosauria'' (second edition). University of California Press:Berkeley, 71-110. . although potentially a valid coelurosaurian. In 2002 Paul Turner found a dorsal vertebra near Grange Chine on the Isle of Wight. An associated partial tibia and metatarsal fragment were subsequently discovered by Oliver Mattsson and referred to ''Calamosaurus''.Naish, D. 2011. Theropod dinosaurs. In Batten, D. J. (ed.) English Wealden Fossils. The Palaeontological Association (London), pp. 526–559. Another specimen referred to ''Calamosaurus'' was collected by local fossil hunter Kai Bailey in 2014. Both specimens are on display at the Dinosaur Expeditions, Conservation and Palaeoart Centre near Brighstone, Isle of Wight. A neck vertebra of a ''Calamosaurus'' was found near Chilton Chine on the Isle of Wight by local fossil hunter Dave Badman. The newly discovered vertebra has gone on display at the Dinosaur Isle Museum in Sandown, Isle of Wight.


Paleobiology

As a possible basal coelurosaur, ''Calamosaurus'' would have been a small, agile,
biped Bipedalism is a form of terrestrial locomotion where an organism moves by means of its two rear limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from Latin ''bis'' 'double' a ...
al
carnivore A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
. Naish ''et al.'' (2001) estimate the living animal would have been around long, with a small head given the build of the neck vertebrae.


References

{{Taxonbar, from=Q1826727 Prehistoric coelurosaurs Barremian life Early Cretaceous dinosaurs of Europe Cretaceous England Fossils of England Fossil taxa described in 1891 Taxa named by Richard Lydekker