Caladenia Graniticola
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''Caladenia graniticola'', commonly known as the Pingaring spider orchid, is a species of
orchid Orchids are plants that belong to the family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Along with the Asteraceae, they are one of the two largest families of flowering ...
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to the
south-west The points of the compass are a set of horizontal, radially arrayed compass directions (or azimuths) used in navigation and cartography. A compass rose is primarily composed of four cardinal directions—north, east, south, and west—each se ...
of
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
. It has a single, hairy leaf and one or two yellowish-green, red and white flowers which have a greenish-yellow and white labellum with a red tip. It was originally described as ''Caladenia hoffmanii'' subsp. ''graniticola'' but has a slightly different labellum and
column A column or pillar in architecture and structural engineering is a structural element that transmits, through compression, the weight of the structure above to other structural elements below. In other words, a column is a compression member. ...
.


Description

''Caladenia graniticola'' is a terrestrial,
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
,
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
,
herb In general use, herbs are a widely distributed and widespread group of plants, excluding vegetables and other plants consumed for macronutrients, with savory or aromatic properties that are used for flavoring and garnishing food, for medicinal ...
with an underground
tuber Tubers are a type of enlarged structure used as storage organs for nutrients in some plants. They are used for the plant's perennation (survival of the winter or dry months), to provide energy and nutrients for regrowth during the next growing ...
and a single erect, hairy leaf, long and wide. One or two flowers long and wide are borne on a stalk tall. The flowers are yellowish-green, red and white and the
lateral Lateral is a geometric term of location which may refer to: Healthcare *Lateral (anatomy), an anatomical direction *Lateral cricoarytenoid muscle *Lateral release (surgery), a surgical procedure on the side of a kneecap Phonetics *Lateral cons ...
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
s and
petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...
s have narrow, club-like,
glandular In animals, a gland is a group of cells in an animal's body that synthesizes substances (such as hormones) for release into the bloodstream (endocrine gland) or into cavities inside the body or its outer surface (exocrine gland). Structure De ...
tips. The lateral sepals and
petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...
s spread widely and curve downwards. The
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal co ...
sepal is erect, long and wide. The lateral sepals are long and wide, curved so that they sometimes cross each other. The
petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...
s are long and about wide and curve downwards. The labellum is long and wide and greenish-yellow with a red tip. The tip of the labellum turns downward but is not rolled under as in some other caladenias. The sides of the labellum have narrow, erect teeth up to long and there are four rows of deep red calli up to long, along the centre of the labellum. Flowering from late September to October.


Taxonomy and naming

The Pingaring spider orchid was first described in 2001 by
Stephen Hopper Stephen Donald Hopper AC FLS FTSE (born 18 June 1951) is a Western Australian botanist. He graduated in Biology, specialising in conservation biology and vascular plants. Hopper has written eight books, and has over 200 publications to his n ...
and Andrew Phillip Brown from a specimen collected near Pingaring, after a specimen was first discovered by Kathleen White in 1984. It was given the name ''Caladenia hoffmanii'' subsp. ''graniticola'' and the description was published in ''
Nuytsia ''Nuytsia floribunda'' is a hemiparasitic tree found in Western Australia. The species is known locally as moodjar and, more recently, the Christmas tree or Western Australian Christmas tree. The display of intensely bright flowers during the ...
''. In 2007, the same authors raised it to species status. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
(''graniticola'') is derived from the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
word ''graniticus'' meaning 'granite rocks' and ''-cola'' meaning 'dweller', referring to the habitat preference of this species.


Distribution and habitat

Pingaring spider orchid occurs between Kalgarin and Newdegate in the
Esperance Plains Esperance Plains, also known as Eyre Botanical District, is a biogeographic region in southern Western Australia on the south coast between the Avon Wheatbelt and Hampton bioregions, and bordered to the north by the Mallee region. It is a pl ...
and Mallee
biogeographic regions A biogeographic realm or ecozone is the broadest biogeographic division of Earth's land surface, based on distributional patterns of terrestrial organisms. They are subdivided into bioregions, which are further subdivided into ecoregions. De ...
where it grows under tall shrubs on and around
granite Granite () is a coarse-grained (phaneritic) intrusive igneous rock composed mostly of quartz, alkali feldspar, and plagioclase. It forms from magma with a high content of silica and alkali metal oxides that slowly cools and solidifies undergro ...
outcrops.


Conservation

''Caladenia graniticola'' is classified as " Threatened Flora (Declared Rare Flora — Extant)" by the Western Australian Government
Department of Parks and Wildlife The Department of Parks and Wildlife (DPaW) was the department of the Government of Western Australia responsible for managing lands described in the ''Conservation and Land Management Act 1984'' and implementing the state's conservation and e ...
. The main threats to the species are its small population size, inappropriate fire regimes, grazing by rabbits and kangaroos and weed invasion.


References

{{Taxonbar, from=Q15287548 graniticola Orchids of Western Australia Endemic orchids of Australia Plants described in 2001 Endemic flora of Western Australia Taxa named by Stephen Hopper Taxa named by Andrew Phillip Brown